For corresponding references and abstracts nos. 1 - 300 see publications , abstracts I, abstracts II, abstracts III, abstracts IV, abstracts V, abstracts VI, and abstracts VII, respectively.
 

408:
A new species of Dysanabatium Bernhauer and additional records of D. jacobsoni Bernhauer (Coleoptera, Staphylinidae, Paederinae). – Dysanabatium hainanense Peng & Li, sp. n. (Hainan: Wuzhi Shan, Diaoluo Shan) is described and illustrated. Additional records of D. jacobsoni Bernhauer, 1915 are reported. The habitus, the sexual characters, and the distribution of this enormously variable species are illustrated.

409:
On the Staphylinidae of Sardinia, Italy (Insecta: Coleoptera). – Recently collected material from Sardinia, comprising more than 1200 specimens and more than 70 species, is studied. Two species are described and illustrated: Astenus (Eurysunius) wunderlei nov.sp. and Leptusa (Adexiopisalia) meybohmi nov.sp. One synonymy is proposed: Anthobium atrocephalum (Gyllenhal 1827) = A. sardoum (Scheerpeltz 1961) nov.syn. Records of zoogeographic interest are reported for 19 species, among them two new records for Sardinia. The currently known distribution of Oxypoda islandica Kraatz 1857 is mapped.

410:
A revision of Palaearctic Lobrathium. V. Three new species from the Himalaya and China, a new synonymy, and additional records (Coleoptera: Staphylinidae: Paederinae). – Four new species of Lobrathium Mulsant & Rey 1878 are described and illustrated: L. subcaeruleum nov.sp. (North India, Nepal), L. wutaium nov.sp. (China: Shanxi: Wutai Shan), L. flexum nov.sp. (China: Jiangxi: Jinggang Shan), and L. circumclusum nov.sp. (Myanmar). Lobrathium zonale Li, Solodovnikov & Zhou 2013, nov.syn. is placed in synonymy with L. configens Assing 2012. Additional records of thirteen species are presented, among them the first record of the genus from Thailand. Lobrathium is now represented in the Palaearctic region by 135 species and one subspecies, with 21 species recorded from the Himalaya and 40 from mainland China.

411:
A revision of Trisunius IV. Two new species and additional records from China (Coleoptera: Staphylinidae: Paederinae). – Two species of the medonine genus Trisunius Assing 2011 from China are described and illustrated: Trisunius smetanai nov.sp. (Yunnan: Gaoligong Shan) and T. rastratus nov.sp. (Hubei: Daba Shan). Additional records of three species are reported from Yunnan. The East Palaearctic genus now includes 19 species.

412:
A revision of Palaearctic and Oriental Rugilus. IV. Three new species from Nepal and additional records (Coleoptera: Staphylinidae: Paederinae). – Three species of the paederine genus Rugilus Leach 1819 from Nepal are described and illustrated: R. (Rugilus) bibarbatus nov.sp. (W-Nepal: Karnali and Seti provinces); R. (R.) schmidti nov.sp. (Manaslu range); R. (R.) bisinuosus nov.sp. (Manaslu range). Additional records of 17 previously described species are reported from the Palaearctic and Oriental regions, among them three new country records. Including the new species, Rugilus is now represented in the Palaearctic and Oriental regions by 97 species and one subspecies.

413:
On Orsunius II. Eight new species and additional records (Coleoptera: Staphylinidae: Paederinae: Medonina). – Eight species of the Oriental medonine genus Orsunius Assing 2011 are described and illustrated: O. spathulatus nov.sp. (Philippines: Mindanao); O. stimulans nov.sp. (North Vietnam, Thailand); O. latexcisus nov.sp. (South Thailand); O. gladiator nov.sp. (Thailand); O. flavovirgatus nov.sp. (South Thailand); O. granulosus nov.sp. (Laos, Thailand, North Vietnam); O. cuneus nov.sp. (Laos); O. heissi nov.sp. (Malaysia: Kedah). Additional records of two previously named species are reported, among them the first records of O. immsi (Bernhauer 1914) from Laos and Thailand. The distributions of all the species of the genus are mapped. A new key to species and an updated catalogue are provided.

414:
Seven new species and additional records of Stilicoderus and Stiliderus (Coleoptera: Staphylinidae: Paederinae). – Five species of Stilicoderus Sharp 1889 and two of Stiliderus Motschulsky 1858 are described and illustrated: Stilicoderus dilatatus nov.sp. (Taiwan) of the S. japonicus group; Stilicoderus seticollis nov.sp. (Indonesia: Maluku: Morotai), S. sociabilis nov.sp. (Maluku: Morotai), S. brevisetosus nov.sp. (Maluku: Morotai), and S. transversus nov.sp. (Maluku: Halmahera) of the S. hieroglyphicus group; Stiliderus aviformis nov.sp. (NW-Thailand) of the S. cicatricosus group; S. tridentatus nov.sp. (Indonesia: Sulawesi Utara) of the S. brendelli group. Additional records of ten described species of Stilicoderus and of four species of Stiliderus are reported.

415:
On Domene scabripennis ROUGEMONT and its close relatives (Coleoptera: Staphylinidae: Paederinae). – Five species of Domene Fauvel 1873, tentatively attributed to the subgenus Macromene Coiffait 1982, from the East Palaearctic and the northern Oriental regions are (re-) described and illustrated: D. scabripennis Rougemont 1995 (Taiwan), D. alesiana nov.sp. (Taiwan), D. immarginata nov.sp. (China: Yunnan), D. sagittata nov.sp. (North Vietnam), and D. firmicornis nov.sp. (China: Zhejiang). These species are assigned to the newly established D. scabripennis group. The placement of D. firmicornis in this group, however, is doubtful and requires confirmation. A key to species is provided. Their distributions are mapped.

416:
Three new species and a new species group of Medon from China and Taiwan (Coleoptera: Staphylinidae: Paederinae). – Three species of the newly established Medon profundus group are described and illustrated: M. profundus nov.sp. (China: border region between Shaanxi and Chongqing, Sichuan), M. yunnanicus nov.sp. (China: Yunnan), and M. alesi (Taiwan). They are evidently closely related and undoubtedly form a monophyletic group. Their known distributions are mapped. The preoccupied name Medon flavomarginatus Assing 2013 is replaced with M. gilvimarginatus nov.nom. Medon Stephens 1833 is now represented in the Palaearctic region by 81 confirmed species.

417:
A Crataraea species associated with Formica chinensis (Coleoptera: Staphylinidae: Aleocharinae). – Crataraea myrmecophila nov.sp. (China: Qinghai) is described and illustrated. The species was exclusively collected from nests of the ant Formica chinensis Wheeler 1913. It represents the first record of pronounced myrmecophily in the genus and the first record of the genus from the East Palaearctic region east of Middle Asia. Crataraea Thomson 1858 is now represented in the Palaearctic region by five species.

418:
New species and records of Lathrobium from the Palaearctic region, primarily from Nepal (Coleoptera: Staphylinidae: Paederinae). – Eighteen species of Lathrobium Gravenhorst, 1802 from Nepal (16 species) and China (2 species) are described and illustrated: L. semicirculatum sp. n. (E-Nepal; L. nepalense group); L. constrictum sp. n. (E-Nepal; L. discissum group); L. descendens sp. n. (W-Nepal; L. deuvei group); L. ascendens sp. n. (W-Nepal; L. deuvei group); L. rude sp. n. (C-Nepal: Dhaulagiri; L. emodense group); L. latibasale sp. n. (W-Nepal; L. pectinatum group); L. eques sp. n. (W-Nepal; L. pectinatum group); L. equitans sp. n. (C-Nepal: Dhaulagiri; L. pectinatum group); L. abruptum sp. n. (W-Nepal; L. pectinatum group); L. kemum sp. n. (C-Nepal: Dhaulagiri; L. pectinatum group); L. flexaculeatum sp. n. (C-Nepal: Dhaulagiri; L. pectinatum group); L. rectaculeatum sp. n. (C-Nepal: Dhaulagiri; L. pectinatum group); L. curvaculeatum sp. n. (C-Nepal: Dhaulagiri; L. pectinatum group); L. hebes sp. n. (C-Nepal: Dhaulagiri; L. pectinatum group); L. imberbe sp. n. (W-Nepal; L. pectinatum group); L. plicatum sp. n. (C-Nepal: Dhaulagiri; L. pectinatum group); L. glabrimpressum sp. n. (China: Yunnan: Gaoligong Shan; group assignment uncertain); L. smetanai sp. n. (China: Gansu: Dalija Shan; L. lentum group). Based on new material, a redescription and new illustrations of L. aculeatum Coiffait, 1982 are provided. Recently collected material of L. calcaratum Assing, 2012 suggests that the holotype of this species may have been mislabelled and that, consequently, the type locality is erroneous. The species previously assigned to the L. aculeatum group are moved to the L. pectinatum group, and the L. aculeatum group is discarded. Several species of the L. pectinatum group were found to be subject to a sexual dimorphism of body size. Additional records of six named species are reported from the West Caucasus (2 species) and from Nepal (4 species). Including the newly described species, the genus is currently represented in the Himalaya by 68 species in ten species groups, in China by 189 species, and in the Palaearctic region by 536 named species and ten subspecies.

419:
A revision of the species  of Pinobius MacLeay, 1871 of the Oriental, Palaearctic, and Australian Regions (Coleoptera: Staphylinidae: Paederinae). – The species of the dolicaonine genus Pinobius MacLeay, 1871 (Coleoptera: Staphylinidae: Paederinae: Dolicaonina) of the Oriental, Palaearctic, and Australian Regions are revised. In all, 28 species are recognized, among them 14 species new to science: P. baculatus sp.n. (Indonesia: Java, Sumba), P. bifidus sp.n. (North India), P. bihamatus sp.n. (Myanmar), P. brevincisus sp.n. (South Vietnam), P. carinatus sp.n. (India: Maharashtra), P. discrepans sp.n. (Australia), P. extensus sp.n. (Papua New Guinea), P. firmilobatus sp.n. (Thailand, Singapore, Peninsular Malaysia, Sabah), P. formidabilis sp.n. (Malaysia), P. imberbis sp.n. (India: Assam), P. inaequalis sp.n. (Indonesia: Sulawesi), P. linguatus sp.n. (Bangladesh), P. rotundatus sp.n. (India: Assam), P. subaequalis sp.n. (Indonesia: Sulawesi). All the species are (re-)described and illustrated. Pinobius vicinus (Kraatz, 1859), previously a synonym of P. indicus (Kraatz, 1859), is revalidated. The following synonymies are proposed: P. indicus (Kraatz, 1859) = longipennis (Cameron, 1931), syn.n.; P. paricolor (Fauvel, 1878) = similis Last, 1984, syn.n.; P. sparsiventris (Fauvel, 1886) = javanus (Cameron, 1936), syn.n. Pinobius pedatus (Lea, 1923) is tentatively regarded as a synonym of P. alatus (Lea, 1923). Lectotypes are designated for Dolicaon adjacens Cameron, 1937, D. alatus Lea, 1923, D. indicus Kraatz, 1859, D. javanus Cameron, 1936, D. robustus Kraatz, 1859, D. sparsiventris Fauvel, 1886, and D. vicinus Kraatz, 1859. The revision revealed that the previously described species had largely been misinterpreted, suggesting that nearly all literature records other than original descriptions are based on misidentifications. Primarily based on the morphology of the aedeagus, the species are assigned to four species groups. The zoogeography of the genus in the study region is discussed. The Indian Subcontinent, eastwards to Myanmar, is the region with the greatest diversity. The distributions of all the species are mapped. The examined material was largely collected at light sources, partly also sifted from leaf litter, at low to intermediate elevations. Only one of the studied specimens is teneral. A key to species and a checklist of the Pinobius species of the Oriental, Palaearctic, and Australian Regions are provided. The genus now comprises a total of 35 species, ten in the Afrotropical, twenty in the Oriental, and seven in the Australian Region. Only five species, none of them exclusive, have been recorded from the south of the Palaearctic Region (southern Arabian Peninsula, South Himalaya, South China), one of them of Afrotropical and the other four of Oriental affiliations. In an appendix, a dolicaonine species from the Afrotropical Region is described: Jarrigeus verberans sp.n. (Nigeria).

420:
A revision of Nazeris. VI. On the fauna of East Yunnan, China (Coleoptera: Staphylinidae: Paederinae). – A recent field trip to eastern Yunnan (Southwest China), a region from where only one species of Nazeris Fauvel, 1873 had been known, yielded thirteen undescribed Nazeris species, eleven of which are described and illustrated: N. brevilobatus sp. n. (Dongchuan env.), N. virilis sp. n. (Wuding env.), N. constrictus sp. n. (Gejiu env.), N. claviger sp. n. (Gejiu env.), and N. discissus sp. n. (Pingbian env.) of the newly established N. brevilobatus group; N. lamellatus sp. n. (Pingbian env.) of the newly established N. lamellatus group; N. semifissus sp. n. (Pingbian env.) of the newly established N. semifissus group; N. fibulatus sp. n. (Kunming env.), N. bulbosus sp. n. (Kunming env.), N. clavilobatus sp. n. (Dongchuan env.), and N. conicus sp. n. (Xundian env.) of the N. cangicus group. Together with N. zhangi Watanabe & Xiao, 1993, the latter three species form a distinct lineage within the N. cangicus group. Two species remain unnamed, as they are represented exclusively by females. The general distribution of the genus in Yunnan and the individual distributions of the newly described species are mapped. Nazeris now includes a total of described 233 species (plus seven subspecies), 118 of which have been recorded from mainland China. The fauna of Yunnan currently comprises 45 described species.

421:
Three new species and additional records of Echiaster from Asia (Coleoptera: Staphylinidae: Paederinae). – Three species of the paederine genus Echiaster Erichson, 1839 from Asia are described and illustrated: Echiaster bivirgatus nov.sp. (Thailand, North India), E. paulexcisus nov.sp. (Thailand), and E. discrepans nov.sp. (South India). Additional records from Thailand, Taiwan, and China are reported for E. unicolor Bernhauer, 1922 and E. maior Assing, 2013. The genus is now represented in Asia by five species in two lineages, one of them composed of one species (E. discrepans) from South India and the other of the remaining four species. A key to species is provided. The distribution of the genus in Asia is mapped.

422:
A revision of Neosclerus Cameron IV. Two new species and additional records from Taiwan (Coleoptera: Staphylinidae: Paederinae). – Two micropterous species of Neosclerus Cameron, 1924 from Taiwan (Nantou Hsien) are described and illustrated, N. brevispinosus nov.sp. and N. biaculeatus nov.sp. Additional records of N. bifidus Assing, 2011 are reported from Taiwan. Including the new species, the genus now includes 31 species and is represented in Taiwan by nine locally endemic species.

423:
Two new species of Eccoptoglossa from Iran and Kyrgyzstan (Coleoptera: Staphylinidae: Aleocharinae). – Eccoptoglossa amischoides nov.sp. (Iran: Ardabil, Lorestan) and E. kirgisica nov.sp. (Kyrgyzstan: Osh) are described and illustrated. Including the new species, the Palaearctic falagriine genus Eccoptoglossa Luze, 1904 now comprises five species and is distributed from the East Caucasus region across Middle Asia eastwards to northwestern Mongolia. The distributions of the Eccoptoglossa species are mapped and a key to species is provided.

424:
On the genus Crataraea (Coleoptera: Staphylinidae: Aleocharinae: Oxypodini). – Three species previously attributed to the Crataraea Thomson, 1858, subtribe Microglottina, are revised and illustrated. Only one of them, C. solskyi (Eppelsheim, 1888), is retained in the genus. Two species are moved to Aleochara Gravenhorst 1802: Aleochara (Xenochara) rubripennis (Fauvel, 1872), nov.comb., and A. (X.) eremita (Peyerimhoff, 1931), nov.comb. The previous synonym Aleochara rubripes Blatchley, 1910 is revalidated to replace the secondary junior homonym Aleochara rubripennis (Casey, 1906). A lectotype is designated for Crataraea rubripennis Fauvel, 1872. Crataraea now includes three species native in the Palaearctic region, with one species adventive in North America. A key to species is provided.

425:
A revision of Tetartopeus IV. A new species from Turkey, new synonymies, and additional records (Coleoptera: Staphylinidae: Paederinae). – Material of the paederine genus Tetartopeus Czwalina, 1888 from the Palaearctic, Oriental, and Australian regions is examined. Tetartopeus korgei nov.sp. (Turkey: Kars), T. lomnickii (Roubal, 1913), and the previously unknown female secondary sexual characters of T. lentus Ryvkin, 1989 are (re-)described and illustrated. The following synonymies are established: Tetartopeus gracilentus (Kraatz, 1859) = Lathrobium pallipes Sharp, 1889, nov.syn., = L. maculatum Last, 1984, nov.syn., = Lobrathium wui Zheng, 2001, nov.syn. = L. bimaculatum Li, Tang & Zhu, 2007, nov.syn. The remarkably vast distribution of T. gracilentus, a species present in the Oriental (Sri Lanka), East Palaearctic, and Australian regions (New Guinea), is mapped. Additional records of fourteen species are presented, among them a new country record.

426:
On the Staphylinidae of Turkey X. Two new species and additional records (Insecta: Coleoptera). – Recently collected material from Turkey, comprising nearly 700 specimens and approximately 130 species, as well as older material from the Korge collection (Berlin), which is now publicly accessible, is studied. Two species of Paederinae and Staphylininae are described and illustrated: Lobrathium unidentatum nov.sp. (Turkey: Hakkâri) and Atanygnathus pullus nov.sp. (Turkey: Adana; Greece: Pelopónnisos); Israel). Four species are reported from Turkey for the first time. The distributions of the newly described species are mapped.

427:
A new species of Platydomene from North India (Coleoptera: Staphylinidae: Paederinae). – Platydomene shavrini nov.sp. from North India (Uttarakhand), the first Platydomene species to be recorded from the Himalaya, is described and illustrated. The genus Platydomene Ganglbauer, 1895 is now represented in the Palaearctic region by 27 species and three subspecies, of which fourteen species are distributed in the East Palaearctic.

428:
A revision of Palaearctic and Oriental Pseudolathra. IV. New species, new combinations and additional records (Coleoptera: Staphylinidae: Paederinae). – Type and non-type material of Pseudolathra Casey, 1905 from the Palaearctic, Oriental, and Australian regions is revised. Four species from Papua New Guinea are transferred from Lathrobium Gravenhorst, 1802 to Pseudolathra, redescribed, and illustrated: Pseudolathra sepikensis (Last, 1984), nov.comb., P. naviculum (Last, 1984), nov.comb, P. puncta (Last, 1984), nov.comb., and P. cyanea (Last, 1984), nov.comb. Two species are described and illustrated: P. armata nov.sp. (Peninsular Malaysia) and P. bisinuata nov.sp. (Malaysia: Borneo). Additional records of five species are reported.

429:
Three new species, redescriptions, a new synonymy, and additional records of Cypha from the West Palaearctic region (Coleoptera: Staphylinidae: Aleocharinae). – Five species of Cypha Leach, 1819 from the West Palaearctic region are (re-)described and illustrated: Cypha rubripennis (Pandellé, 1869) (France, Spain, Morocco); C. rubicunda (Reitter, 1887) (Turkmenistan); C. truncata nov.sp. (Ukraine); C. prominens nov.sp. (Tunisia); C. persica nov.sp. (Iran). Cypha tbilisiensis Dauphin, 2005 nov.syn. is placed in synonymy with C. tarsalis (Luze, 1902). Several new country records are reported. The genus is now represented in the Palaearctic region by 50 described species.

430:
On the Staphylinidae of Israel II, with a revision of some species of Dinusa Saulcy (Coleoptera: Staphylinidae: Aleocharinae). – The results of extensive pitfall trap studies conducted in Israel in 2010 and 2011 are presented, and material of some species of Dinusa Saulcy, 1865 is revised. Records of zoogeographic interest of 49 species are reported, among them 25 first records from Israel, three from Jordan, three from Iraq, one from Syria, one from Egypt, and one from Malta. Three species of Paederinae and Aleocharinae from Israel and one species of Aleocharinae from southern Turkey are described and illustrated: Astenus (Eurysunius) affimbriatus nov.sp. (Israel: Hula valley), Pseudosemiris torta nov.sp. (Israel: numerous localities), Geostiba (Sibiota) meronica nov.sp. (Israel: Mount Meron), and Dinusa subangulata nov.sp. (Turkey: Gaziantep). The distributions of Astenus affimbriatus and Pseudosemiris torta are mapped. Astenus (Eurysunius) platynotus (Saulcy, 1865) is redescribed and illustrated. Neotypes are designated for Sunius platynotus Saulcy, 1865 and Dinusa jebusaea Saulcy, 1865. Lectotypes are designated for Dinusa hierosolymata Saulcy, 1865, D. particeps Abeille de Perrin, 1896, and D. angulicollis Fauvel, 1886. The following taxonomic and nomenclatural acts are proposed: Quedius josue Saulcy, 1865 = Q. troodites Fagel, 1968, nov.syn.; Dinusa hierosolymata Saulcy, 1865 = D. saulcyi Abeille de Perrin, 1896, nov.syn.; Dinusa jebusaea Saulcy, 1865 = D. particeps Abeille de Perrin, 1896, nov.syn.; Notothecta hipponensis (Fauvel, 1886), nov.comb. (ex Dinusa). Dinusa santschii Wasmann, 1912 from Tunisia is excluded from Dinusa and tentatively moved to Cypha Leach, 1819. Dinusa now contains eight myrmecophilous species associated with harvester ants (Messor spp.) and distributed in the East Mediterranean from southern Greece across Turkey to the Middle East. A checklist of the named species of Dinusa is provided.

431:
A revision of Pronomaea Erichson. II. A new species from China and additional records (Coleoptera: Staphylinidae: Aleocharinae). – Additional records of four species of Pronomaea Erichson, 1837 are reported from the West Palaearctic region, among them the first record of P. araxicola Reitter, 1898 from Afghanistan. Pronomaea maxima nov.sp. from Yunnan (China) is described and illustrated. The genus is now represented in the Palaearctic region by seventeen species.

432:
A revision of the Habrocerinae of the world. V. A new species of Habrocerus from  China and additional records (Coleoptera: Staphylinidae). – Habrocerus bisetosus nov.sp. (China: S-Gansu: Qinling Shan) is described, illustrated, and distinguished from other species of the H. capillaricornis group. Additional records of seven named and one unnamed species of Habrocerus Erichson, 1839 are reported from the Palaearctic region and Central America (Guatemala), respectively, among them the remarkable record of a male of H. indicus Assing & Wunderle, 1995 from southeastern Yunnan, China. Its sexual characters are illustrated. The distributions of H. indicus in the East Palaearctic region and of Habrocerus in China are mapped. Habrocerus is now represented by 17 and the Habrocerinae by 23 named species worldwide. An updated world catalogue of Habrocerinae is provided.

433:
On the Oedichirus fauna of China (Coleoptera: Staphylinidae: Paederinae). – Three micropterous species of Oedichirus Erichson, 1839 from Yunnan, China, are described and illustrated: O. latexcisus nov.sp. (environs of Dongchuan), O. schuelkei nov.sp. (Diancang Shan), and O. abbreviatus nov.sp. (environs of Tengchong). The genus is now represented in China and Taiwan by six described species, four of them micropterous. A key to the species of China and Taiwan is provided. The distributions of the micropterous species are mapped.

434:
A revision of the Megalopaederus species of Taiwan (Coleoptera: Staphylinidae: Paederinae). – Megalopaederus Scheerpeltz, 1957, previously regarded as a distinct genus, is assigned to the genus Paederus Fabricius, 1775 as a subgenus. The Megalopaederus species of Taiwan are revised. In all, eighteen named locally endemic species belonging to two species groups are recognized, thirteen of them are described and illustrated for the first time: Paederus (Megalopaederus) virgifer n. sp. (Nantou, Chiai); P. (M.) incurvatus n. sp. (Kaohsiung); P. (M.) adiectus n. sp. (Pingtung); P. (M.) aequilobatus n. sp. (Pingtung); P. (M.) furcispinosus n. sp. (Nantou); P. (M.) excisissimus n. sp. (Pingtung); P. (M.) trihamatus n. sp. (Taoyuan); P. (M.) bivirgatus n. sp. (Hualien); P. (M.) anmamontis n. sp. (Taichung); P. (M.) semiflavus n. sp. (Taitung); P. (M.) alesi n. sp. (Hualien); P. (M.) discurrens n. sp. (Taichung); P. (M.) incisus n. sp. (Nantou). Rescriptions and illustrations of the five previously described species, as well as of P. (M.) lewisi Cameron, 1930 from Japan and P. biacutus Li et al., 2013 from mainland China are provided, and P. (M.) bucculentus n. sp. (Japan: Honshu: Nagano Prefecture) is described. The latter three species were included in the revision to tentatively assess the phylogenetic affiliations of Taiwanese Megalopaederus. Paederus fulvocaudatus Adachi, 1939, previously considered a subspecies of P. kosempoensis Bernhauer, 1922, represents a distinct species. A lectotype is designated for P. kosempoensis Bernhauer, 1922. Additional material composed exclusively of females and probably representing six undescribed species is listed. The available information on the natural history of Taiwanese Megalopaederus is compiled. A key to the named species of Taiwan is provided. Their distributions are mapped.

435:
On the Staphylinidae (Coleoptera) of Crete II. Seven new species, a new synonymy, and additional records. – Approximately 110 species of Staphylinidae collected in 2013 and 2014 are reported from the Greek island Crete. Seven species of Aleocharinae (four species), Paederinae (one species), and Staphylininae (two species) are described and illustrated, five of them are probably local endemics: Bellatheta albimontis n. sp. (Lefka Ori); B. idana n. sp. (Psiloritis); Geostiba (Sipalotricha) paulexcisa sp n. (Psiloritis); Myrmecopora (Myrmecopora) thriptica n. sp. (Orno Thriptis; associated with Messor sp.); Sunius thripticus n. sp. (Orno Thriptis); Quedius praecisus n. sp. (Orno Thriptis, but probably more widespread in Crete); Xantholinus erinaceus n. sp. (several localities). The sexual characters of some previously described species are illustrated. The following synonymy is proposed: Quedius nemoralis Baudi di Selve, 1848 = Q. candicus Coiffait, 1976, syn. n. The genus Bellatheta Roubal, 1928 is reported from Crete for the first time. Some previous records are rectified. The diversity and biogeography of the endemic species of Crete is discussed. A revised list of the 67 currently known named endemic (sub-)species of Staphylinidae of Crete is provided.

436:
Three new species of Domene from China (Coleoptera: Staphylinidae: Paederinae). – Three species of Domene Fauvel, 1873 from China, are described and illustrated: Domene (Macromene) affimbriata sp. n. (southeastern Yunnan: Dawei Shan near Pingbian), D. (M.) praefigens sp. n. (eastern Yunnan: region to the north, northwest, and northeast of Kunming), and D. (M.) emeiana sp. n. (Sichuan: Emei Shan). The distributions of the new species and other named species of the genus in China and Taiwan are mapped. Domene now includes 64 named species, 15 of which have been recorded from China (13) and Taiwan (2).

437:
New species and additional records of Lathrobium and Elytrobium from the Palaearctic region, with special reference to the fauna of East Yunnan (Coleoptera: Staphylinidae: Paederinae). – Seventeen species of Lathrobium Gravenhorst, 1802 and one of Elytrobium Assing, 2013 are (re-)described and illustrated: Lathrobium struyvei sp. n. (Spain), a close relative of L. dimidiatipenne Bernhauer, 1910; L. kiruense sp. n. (Nepal: Annapurna) of the L. emodense group; L. undosum sp. n. (Nepal: Solu Khumbu), tentatively assigned to the L. discissum group; L. crenatum sp. n. (China: East Yunnan), L. rostratum sp. n. (China: East Yunnan), L. tricarinatum sp. n. (China: East Yunnan), and L. laciniatum sp. n. (China: East Yunnan), all four of them of the L. tentaculatum group; L. daweianum sp. n. (China: Southeast Yunnan) of the newly established L. daweianum group; L. coadultum sp. n. (China: Southeast Yunnan) of the newly established L. coadultum group; L. grebennikovi sp. n. (China: Shaanxi: Qinling Shan) and L. cavisulcatum sp. n. (China: Shaanxi: Qinling Shan) of the L. varisternale group; L. abruptum sp. n. (China: Sichuan: Gongga Shan), a close relative of L. hailuogouense Peng et al., 2012; L. erectum sp. n. (China: Guizhou: Fanjing Shan) of the L. fissispinosum group; L. scaphiforme sp. n. (Japan: Honshu); L. volutum sp. n. (Japan: Honshu); L. trabale sp. n. (Japan: Honshu); L. oharai Watanabe, 2014; Elytrobium edentulum sp. n. (China: Shaanxi: Qinling Shan). The diversity, biogeography, natural history, and phylogenetic affiliations of the Lathrobium fauna of East Yunnan, from where the genus is reported for the first time, are discussed. The general distribution of Lathrobium and the individual distributions of the newly described species in Yunnan are mapped. Additional records of 30 described and one unnamed Lathrobium species are reported from Middle Asia (one species), Nepal (five species), from the Chinese provinces Yunnan (five species), Sichuan (five species), Shaanxi (one species), and Guizhou (one species), from Taiwan (one species), and from Japan (13 species). Three of these species are reported from Japan for the first time. Additional records of two Elytrobium species are reported from China and Japan. Lathrobium is now represented in the Palaearctic region by 563 species and ten subspecies, in China by 206 species (67 of which are confined to Yunnan), in Japan by 114, and in the Himalaya by 70 species. Elytrobium currently includes a total of seven species distributed in China, Taiwan, and Japan.

438:
On the Staphylinidae of the Greek island Chios (Insecta: Coleoptera). – A study of 536 specimens of Staphylinidae collected in the Greek island Chios in December 2014 yielded 41 species, among them at least three undescribed island-endemic species and two first records from Greece. A species list is provided. Xantholinus (Helicophallus) chiosicus nov.sp., at present the sole named island-endemic representative of the family in Chios, is described and illustrated. Geostiba bulbifera Zerche, 1988, nov.syn. is placed in synonymy with G. maxiana (Tikhomirova, 1973). The first discovery of a winged male of Geostiba maxiana confirms an earlier hypothesis that this species is wing-dimorphic.

439:
A new species of Phloeocharis Mannerheim from Spain, with a note on P. brachyptera Sharp (Coleoptera: Staphylinidae: Phloeocharinae). – Phloeocharis gredosensis nov.sp. (Central Spain: Sierra de Gredos) is described, illustrated, and distinguished from other micropterous Phloeocharis species known from Spain. Material of P. brachyptera Sharp, 1873 from various regions in the Cordillera Cantabria and the western Pyrenees (northern Spain) is studied. Based on an examination of the male sexual characters, the previously established synonymy of P. umbratilis Eppelsheim, 1880 with P. brachyptera is confirmed. The male sexual characters of P. brachyptera and P. microptera Fauvel, 1898 are illustrated. Phloeocharis Mannerheim, 1830 now includes 45 species, twelve of which have been recorded from Spain.

440:
A revision of Amarochara Thomson of the Holarctic region V. A new species from China, a new combination, the male of A. caeca Assing, and additional records (Coleoptera: Staphylinidae: Aleocharinae: Aleocharini). – Amarochara daweiana nov.sp. (China: Yunnan) is described and illustrated. Additional records of species of Amarochara Thomson, 1858 are reported from the Palaearctic region, among them several first records from China and Laos. The previously unknown male sexual characters of A. caeca Assing, 2002 are described and illustrated. A new combination is established: Ocalea nepalica (Pace, 2013) nov.comb. (ex Amarochara). The genus is now represented in the Holarctic region by 31 species.

441:
A revision of Palaearctic and Oriental Rugilus. V. Two new species from China and additional records (Coleoptera: Staphylinidae: Paederinae). – Two species of Rugilus Leach, 1819 from China are described and illustrated: R. (Rugilus) emeimontis nov.sp. (Sichuan: Emei Shan), the third endemic Rugilus species from the Emei Shan, and R. (R.) rugosissimus nov.sp. (Guizhou: Fanjing Shan), the first record of Rugilus sensu strictu from Guizhou. Additional records of 15 previously described species are reported from the Palaearctic and Oriental regions, among them the first record of R. dabaicus Assing, 2012 from the Qinling Shan (Shaanxi). Two probably undescribed species from Yunnan and Guizhou, which are currently represented only by females, remain unnamed. Including the new species, Rugilus is now represented in the Palaearctic and Oriental regions by 100 species and one subspecies.

442:
On Orsunius III. Four new species from China and Thailand, and additional records (Coleoptera: Staphylinidae: Paederinae: Medonina). – Four species of Orsunius Assing, 2011 are described and illustrated: O. affimbriatus nov.sp. (China: Guangdong); O. incurvatus nov.sp. (Thailand); O. granulosissimus nov.sp. (China: Yunnan); O. confluens nov.sp. (China: Yunnan). Additional records of four previously described species are reported, among them the first record of O. cuneatus Assing, 2014 from Thailand. A new key to species and an updated catalogue are provided. The genus currently comprises 21 named species.

443:
On the identity of Atheta opacicollis (Fauvel) (Coleoptera: Staphylinidae: Aleocharinae: Oxypodini). – Types and additional material of Atheta opacicollis (Fauvel, 1878) are revised. A lectotype is designated. The species is reported from Syria for the first time. Diagnostic characters distinguishing A. opacicollis from the similar A. meybohmi Assing, 2011 are described and illustrated.

444:
On the Orphnebius fauna of the East Palaearctic region. VI. Six new species from China and Taiwan, and additional records (Coleoptera: Staphylinidae: Aleocharinae: Lomechusini). – Six species of Orphnebius Motschulsky, 1858 are described and illustrated: Orphnebius dishamatus nov.sp. (Yunnan), O. incrassatus nov.sp. (Yunnan), and O. formosanus nov.sp. (Taiwan) of the O. hauseri group; O. tridentatus nov.sp. (Yunnan) of the O. nanlingensis group; O. (Deroleptus) multimpressus nov.sp. (Yunnan); O. (Deroleptus) planicollis nov.sp. (Yunnan). Deroleptus Bernhauer, 1915, previously regarded as a distinct genus, is placed in Orphnebius as a subgenus. Additional records of nine species from North India, Nepal, and China are reported. An updated catalogue of the Orphnebius fauna of the East Palaearctic region and a revised key to the species of China and Taiwan are provided. Orphnebius is now represented in the East Palaearctic region by 38 named species, 19 of which are distributed in the Himalaya, 18 in China, and one in Taiwan.

445:
On the Tetrabothrus fauna of China (Coleoptera: Staphylinidae: Aleocharinae: Lomechusini). – Five species of Tetrabothrus Bernhauer, 1915 from China are (re-) described and illustrated: T. cavus nov.sp. (Yunnan, Shaanxi), T. inflexus nov.sp. (Yunnan), T. brevalatus nov.sp. (Yunnan), T. rubricollis nov.sp. (North Sichuan), and T. chinensis Pace, 2012. The six named species recorded from China thus far are keyed and their distributions are mapped. In total, the Tetrabothrus material known from China at present amounts to only twelve specimens, one of them unnamed. Including the new species, the genus currently includes 28 species, 18 of which have been recorded from the southern East Palaearctic region.

446:
On Stilicoderus and Stiliderus IV. Two new species from China and additional records (Coleoptera: Staphylinidae: Paederinae). – Stilicoderus hainanus nov.sp., the first representative of the genus to be recorded from the Chinese island Hainan, and S. daweianus nov.sp. (China: Yunnan: Dawei Shan) are described and illustrated. Additional records of eight previously described species are reported. The distributions of four species in China are mapped. An updated catalogue of the species of Stilicoderus Sharp, 1889 and Stiliderus Motschulsky, 1858 recorded from the East Palaearctic region, including Northeast India and Burma, is provided. Stilicoderus now comprises 107 species, 39 of which have been recorded from the East Palaearctic region. The country with, by far, the most diverse Stilicoderus fauna is China (26 species), where as many as 18 species have been recorded from Yunnan alone.

447:
On the Harpopaederus fauna of China (Coleoptera: Staphylinidae: Paederinae). – Types and additional material of Paederus Fabricius, 1775, subgenus Harpopaederus Scheerpeltz, 1957, from China are revised. Seven species are (re-) described and/or illustrated: Paederus (Harpopaederus) dangchangensis Li & Zhou, 2007; P. (H.) lineodenticulatus Li & Zhou, 2007; P. (H.) cultellatus nov.sp. (Daba Shan at the border between Chongqing, Hubei, and Shaanxi); P. (H.) deplectens nov.sp. (Northwest Sichuan); P. (H.) edentulus nov.sp. (Gansu: mountains to the southeast of Longnan); P. (H.) minicus nov.sp. (Gansu: Min Shan); P. (H.) chinensis Bernhauer, 1931. Paederus chinensis, the type species of the subgenus Oedopaederus, is assigned to Harpopaederus, resulting in the synonymy Harpopaederus Scheerpeltz, 1957 = Oedopaederus Scheerpeltz, 1957, nov.syn. Paederus pseudobaudii Aleksandrov, 1934, nov.syn., is placed in synonymy with P. gottschei Kolbe, 1886. The intraspecific variation of the sexual characters in P. dangchangensis is illustrated and discussed. Additional records of nine previously described species are reported. The general distribution of the subgenus in China, as well as the individual distributions of all the Harpopaederus species recorded from China are illustrated. A catalogue is provided. Harpopaederus now includes 20 species, 15 of which have been recorded from China (14 exclusive). With one exception (P. gottschei), all the Chinese representatives of the subgenus are distributed in the Qinling Shan, the Daba Shan, and adjacent mountain ranges, as well as in West Sichuan.

448:
Two new genera of Medonina from China and Japan (Coleoptera: Staphylinidae: Paederinae). – Granimedon nov.gen. (type species: G. anguliceps nov.sp.) and Coimedon nov.gen. (type species: Lithocharis dissimilis Sharp, 1874) are described and illustrated. The former includes three species, all of them from southeastern Yunnan, China: Granimedon anguliceps nov.sp.; G. effeminatus nov.sp.; G. creber nov.sp. Coimedon is represented only by C. dissimilis (Sharp, 1874), nov.comb., from Kyushu, South Japan. All four species are (re-)described and illustrated. A key to the species of Granimedon is provided. The distributions of the Granimedon species are mapped.

449:
On the Nepalota fauna of China (Coleoptera: Staphylinidae: Aleocharinae: Athetini). – Types and additonal material of the East Palaearctic aleocharine genus Nepalota Pace, 1987 are examined. Eleven species are (re-)described and illustrated: N. franzi Pace, 1987; N. globifera Pace, 1998; N. smetanai Pace, 1998; N. guangdongensis Pace, 2004; N. fellowesi Pace, 2004; N. crocea nov.sp. (Yunnan); N. cuneata nov.sp. (Yunnan); N. mocytoides nov.sp. (East Yunnan); N. prominula nov.sp. (Yunnan); N. tuberifera nov.sp. (Yunnan); N. daweiana nov.sp. (Yunnan). Five synonymies are proposed: N. gansuensis Pace, 1998 = N. qinlingmontis Pace, 2011, nov.syn.; N. globifera Pace, 1998 = N. caluoensis Pace, 2011, nov.syn., = N. daxuensis Pace, 2011, nov.syn.; N. guangdongensis Pace, 2004 = N. rougemonti Pace, 2011, nov.syn. = N. ruficollis Pace, 2011, nov.syn. Nepalota franzi is reported from China for the first time. In all, 35 valid species are currently known from the East Palaearctic, four of them of doubtful status (males unknown). Twenty species have been described or reliably recorded from China exclusive of Taiwan. The diversity is greatest in Yunnan (fourteen species, seven of them exclusive), followed by Hubei (five species; three exclusive); Sichuan (four species, one exclusive), Gansu and Shaanxi (three species each), Guangdong (two species), Zhejiang, Chongqing, Jiangxi, Guangxi, Guizhou, and Beijing (one species each). The distributions of 15 species are mapped. An updated checklist of Nepalota is provided.

450:
Two new species of Chinecallicerus (Coleoptera: Staphylinidae: Aleocharinae: Geostibini). – Chinecallicerus subater nov.sp. (China: Northeast Yunnan) and C. reticulatus nov.sp. (China: Southeast Yunnan) are described, illustrated, and distinguished from their congeners. An updated key to the species of Chinecallicerus Assing, 2004, a genus currently known only from Yunnan, China, and including six species is provided. The distributions of the genus and the individual species are mapped.

451:
A revision of Porocallus Sharp. IV. A new species from China and additional records (Coleoptera: Staphylinidae: Aleocharinae). – Porocallus cicatricatus nov.sp. (China: southeastern Yunnan) is described and illustrated. Additional records of two previously described species are reported. The currently known distributions of all the species of Porocallus Sharp, 1888 are mapped. The East Palaearctic genus now includes seven species.

452:
A revision of Nazeris VII. Three new species and additional records from Nepal and China (Coleoptera: Staphylinidae: Paederinae). – Three species of Nazeris Fauvel, 1873 are described and illustrated: N. brevior sp. n. (Nepal: Solu Khumbu) of the N. alticola group; N. hamulatus sp. n. (China: Shaanxi: Qinling Shan) of the N. longilobatus group; N. eminens sp. n. (China: Sichuan: Emei Shan), tentatively assigned to the N. zhangi lineage of the N. cangicus group. Additional records of eleven described and one unnamed species are reported from Nepal (four species) and China (eight species). The Palaearctic genus is now represented by 236 described species (plus seven subspecies), 120 of which are distributed in mainland China and 49 in the Himalaya.

453:
On the Staphylinidae of the Greek island Samos (Coleoptera: Staphylinidae). – A recent field trip to the Greek island Samos yielded 1322 specimens belonging to at least 117 species of Staphylinidae. Three species are described and illustrated: Geostiba (Tropogastrosipalia) plicipennis sp. n. (Oros Ambelos) and Oxypoda (Bessopora) kerkisica sp. n. (Oros Kerkis) of the Aleocharinae, and Sunius ambelosicus sp. n. (Oros Ambelos) of the Paederinae. Geostiba plicipennis is the first representative of the speciose subgenus Tropogastrosipalia Scheerpeltz, 1951 to be recorded from an island. Including the Pselaphinae and Scydmaeninae recorded by Volker Brachat and Heinrich Meybohm in 2003, as well as previous literature records, at least 136 species of Staphylinidae are currently known from Samos, 103 of them named and positively identified, 15 undescribed (five of the Pselaphinae and ten of the Scydmaeninae), eight tentatively identified, and at least ten unidentified. Six species are reported from Greece for the first time. The remarkably diverse staphylinid fauna of Samos includes as many as 18 species that have been recorded exclusively from this island and are most likely island-endemic, some of them even locally endemic. However, only seven of these species have been named; the remainder is pending description. Xantholinus chiosicus Assing, 2015, a species previously hypothesized to be endemic to Chios, is recorded also from Samos. The diversity of the staphylinid fauna of Samos is evidently greater than that of other North Aegean islands; it appears to be even greater than that of the much larger island Rhodes, an observation that may be explained with the geography, geology, and ecological characteristics of Samos. A comprehensive list of the named and unnamed species currently known from Samos is provided.

454:
New species and additional records of Lomechusini from the Palaearctic region, primarily from China (Coleoptera: Staphylinidae: Aleocharinae). – Six species of Lomechusini from China and North India are (re-)described and illustrated: Drusilla yunnanensis Pace, 1993; D. watanabei Maruyama & Kishimoto, 2002; D. flagellata sp. n. (China: Yunnan); D. palata sp. n. (China: Yunnan); Amaurodera angularis sp. n. (North India: West Bengal); Lomechusoides penicillatus sp. n. (China: Gansu). Two synonymies are proposed: Drusilla canaliculata (Fabricius, 1787) = D. puncticollis Motschulsky, 1845, syn. n.; Amaurodera smetanai Pace, 1992 = A. yaoana Pace, 1992, syn. n. Additional records of 29 previously described species of Drusilla Leach, 1819, Pella Stephens, 1835, Amaurodera Fauvel, 1905, Tetrabothrus Bernhauer, 1915, and Lomechusoides Tottenham, 1939 are reported from the Palaearctic region, primarily from China, among them several new country records and the first record of Pella lugens (Gravenhorst, 1802) from the East Palaearctic. The previously unknown aedeagus of Pella reuteri Assing, 2008, the previously unknown female sexual characters of Amaurodera ilamica Assing, 2005, and the primary sexual characters of several Pella species recorded from China are illustrated. The distributions of the Drusilla species currently known from China are illustrated.

455:
A revision of Neosclerus V. Thirteen new species and additional records (Coleoptera: Staphylinidae: Paederinae). – Thirteen species of Neosclerus Cameron, 1924 are described and illustrated: N. albens sp. n. (Sri Lanka), N. areolatus sp. n. (Thailand), N. biangulatus sp. n. (Thailand), N. daweianus sp. n. (China: Yunnan), N. discolor sp. n. (Burma, Thailand), N. figens sp. n. (China: Yunnan), N. forficatus sp. n. (Sri Lanka), N. fractus sp. n. (India: Meghalaya), N. khasicus sp. n. (India: Meghalaya), N. revolutus sp. n. (South India), N. sagittatus sp. n. (India: Meghalaya), N. semicalvus sp. n. (Thailand), and N. trisinuatus sp. n. (China: Yunnan). The previously unknown male sexual characters of N. granulicollis Cameron, 1924 are described and illustrated. Additional records of twelve named and three unnamed species are reported, among them several new country records. The genus now comprises 44 named species in five species groups, one of them newly established. The countries with the greatest diversity are China, India, and Thailand (ten species each), and Taiwan (nine locally endemic species). The distribution of the genus as a whole, as well as the individual distributions of 34 species are mapped. An updated catalogue is provided.

456:
A revision of Sunius XV. Six new species from Kyrgyzstan and the Himalaya, and additional records (Coleoptera: Staphylinidae: Paederinae). – Six micropterous species of Sunius Stephens, 1829 are described and illustrated: S. alaianus sp. n. (Kyrgyzstan) of the S. viator group; S. marratus sp. n. (North India: West Bengal), S. virilis (North India: West Bengal), S. deplectens sp. n. (North India: West Bengal), S. dearmatus sp. n. (East Nepal), and S. migrus sp. n. (Central Nepal), all five of them of the S. manasluensis group. Additional records of 16 species are reported both from the West and the East Palaearctic regions. The genus is now represented in the Palaearctic region by 126 described species and two subspecies.

457:
The first species of Borboropora KRAATZ from the East Palaearctic region (Coleoptera: Staphylinidae: Aleocharinae: Falagriini). – Borboropora indica nov.sp. (India: Arunachal Pradesh), the first representative of the genus recorded from the East Palaearctic region, is described and illustrated. Borboropora pseudoquadriceps Delgado & Santiago-Jiménez, 2009 and B. mixe Delgado & Santiago-Jiménez, 2009 from Mexico are excluded from Borboropora Kraatz, 1862 and tentatively moved to Anaulacaspis Ganglbauer, 1895 and Falagrioma Casey, 1906, respectively, resulting in the binomina Anaulacaspis pseudoquadriceps (Delgado & Santiago-Jiménez, 2009), nov.comb., and Falagrioma mixe (Delgado & Santiago-Jiménez, 2009), nov.comb. Thus, Borboropora currrently includes a total of seven species..

458:
A new myrmecophilous species of Eurysunius from Turkey (Coleoptera: Staphylinidae: Paederinae). – Astenus (Eurysunius) kociani nov.sp. (Turkey: Nevşehir) is described, illustrated, and distinguished from other species of Eurysunius Reitter, 1909 known from Turkey, particularly the similar and evidently closely allied Astenus sexsetosus Assing, 2002. The species was found associated with an unidentified species of Tetramorium Mayr, 1855.

459:
New species and additional records of Dolicaonina from Ecuador (Coleoptera: Staphylinidae: Paederinae). – Five species of Gnathymenus Solier, 1849 are described and illustrated, all of them from Ecuador: Gnathymenus penicillatus nov.sp. (Cotopaxi); G. crucifer nov.sp. (Zamora Chinchipe); G. stipulatus nov.sp. (Zamora Chinchipe); G. abscisus nov.sp. (Zamora Chinchipe); G. multiplicus nov.sp. (Esmeraldas). One of these species (G. penicillatus) is micropterous, the remainder macropterous. Additional records of three species of Gnathymenus and one of Stenopholea Herman, 1969 are reported from Ecuador.

460:
A revision of the Lithocharis species of the Palaearctic, Oriental and Australian regions (Coleoptera: Staphylinidae: Paederinae: Medonina). – The species of the medonine genus Lithocharis Dejean, 1833 of the Palaearctic (Mauritania included), Oriental, and Australian regions are revised. In all, 23 species are recognized, one of them of doubtful identity (male and type depository unknown) and one (male unknown) of doubtful generic assignment. Except for few, partly previously revised species from the West Palaearctic region, all the species are (re-)described and illustrated, six of them for the first time: Lithocharis socotrana nov.sp. (Yemen: Socotra); L. bitriangulata nov.sp. (Sumatra); L. inermis nov.sp. (Thailand); L. latexcisa nov.sp. (Nepal); L. ornatrix nov.sp. (Malaysia, Indonesia); L. parvincisa nov.sp. (Indonesia: Sulawesi). Lithocharis erythroptera Gemminger & Harold, 1868 is revalidated. Five synonymies are established: L. erythroptera Gemminger & Harold, 1868 = L. penicillata Cameron, 1928, nov.syn.; L. nigriceps Kraatz, 1859 = L. changlingensis Li, 1992, nov.syn.; L. vilis Kraatz, 1859 = L. jacobsoni Cameron, 1928, nov.syn., = L. vexans Cameron, 1936, nov.syn.; Panscopaeus lithocharoides (Sharp, 1874) = Medon subopacus Bernhauer, 1907, nov.syn. Lithocharis gigantea Fauvel, 1878 is excluded from Lithocharis. Lithocharis ochracea (Gravenhorst, 1802) is confirmed as a valid name and removed from the previously proposed synonymy with Staphylinus tricolor Fabricius, 1787, whose valid present combination is Xantholinus tricolor (Fabricius, 1787). Lectotypes are designated for Lithocharis sororcula Kraatz, 1859, L. fuscipennis Kraatz, 1859, L. penicillata Cameron, 1928, L. uvida Kraatz, 1859, L. lamellifera Cameron, 1928, L. distinguenda Cameron, 1928, L. sordida Cameron, 1928, L. timorensis Cameron, 1928, Medon subopacus Bernhauer, 1907, and Staphylinus tricolor Fabricius, 1787. Inhabitants of decaying organic matter and active flyers, Lithocharis species are generally remarkably widespread. Males of this genus are occasionally subject to conspicuous teratologies (aedeagus completely reduced; male secondary characters absent). A key to the Lithocharis species of the Palaearctic (including Mauritania), Oriental, and Australian regions, and a catalogue are provided.

461:
A revision of Trisunius V. Three new species and additional records, including the first records of the genus from Vietnam (Coleoptera: Staphylinidae: Paederinae). – Three species of the medonine genus Trisunius Assing, 2011 from North Thailand are described and illustrated: Trisunius scaphiformis nov.sp. (Doi Inthanon), T. conlectus nov.sp. (Doi Inthanon), and T. penicillatus nov.sp. (Doi Pha Hom). Additional records of four species are reported from East Yunnan, Thailand, and Vietnam. The discovery of two species in Vietnam represents the first record of the Trisunius from this country. The distribution of the genus as a whole and the individual distributions of three widespread species are mapped. Trisunius now includes 22 species.

462:
First record of Pseudomedon from the Oriental Region (Coleoptera: Staphylinidae: Paederinae: Medonina). – Pseudomedon discolor nov.sp. (North Vietnam: environs of Sa Pa), representing the first record of the genus from the Oriental region and from Vietnam, is described and illustrated. An additional record of Pseudomedon kazakhstanicus Assing, 2008 is reported from Kazakhstan. The genus now includes a total of 23 species.

463:
A revision of Sclerochiton. II. Two new species and additional records, including the first records of the genus from Vietnam (Coleoptera: Staphylinidae: Paederinae). – Two species of Sclerochiton Kraatz, 1859 are described and illustrated: Sclerochiton pectinatus nov.sp. (Laos) and S. peniculatus nov.sp. (North Vietnam). Additional records of ten species are reported. The genus is recorded from Vietnam for the first time (two species). The distribution of the genus in the southern East Palaearctic and Oriental regions and the individual distributions of all the species treated are mapped. Sclerochiton now includes a total of 23 described species, 17 of which are distributed in the southern East Palaearctic and Oriental regions.

464:
A revision of the Habrocerinae of the world. VI. The first records of the subfamily from Vietnam (Coleoptera: Staphylinidae). – Habrocerus wunderlei nov.sp. (North Vietnam: environs of Sa Pa) is described, illustrated, and distinguished from other species of the H. schwarzi group. Additional records of two species are reported from North Vietnam and Thailand. The subfamily Habrocerinae is reported from Vietnam for the first time. Habrocerus Erichson, 1839 is now represented by 18 and the Habrocerinae by 24 named species worldwide. A revised and updated world catalogue of subfamily is provided.

465:
Three new species of Deinopteroloma from Vietnam and China (Coleoptera: Staphylinidae: Omaliinae). – Three species of Deinopteroloma Jansson, 1946 are described and illustrated: Deinopteroloma uncinatum nov.sp. (North Vietnam: environs of Sa Pa), D. dispar nov.sp. (North Vietnam: environs of Sa Pa), and D. obtortum nov.sp. (China: southern Gansu). The genus is reported from Vietnam and the Oriental region for the first time. Additional records of D. chiangi Smetana, 1990 and D. gracile Smetana, 2001 are reported from Taiwan and Sichuan, respectively. The distribution of the genus in Asia east of the Himalaya is mapped. Including the new species, Deinopteroloma currently includes 17 species and is distributed in the western Nearctic, the southern East Palaearctic, and the northern Oriental regions.

466:
A revision of Porocallus SHARP. V. The first records of the genus from Vietnam (Coleoptera: Staphylinidae: Aleocharinae). – Porocallus vietnamensis nov.sp. (North Vietnam: environs of Sa Pa) is described and illustrated. Porocallus wrasei Assing, 2012, a species previously known only from the Chinese province Yunnan, is reported from Vietnam for the first time. The East Palaearctic genus Porocallus Sharp, 1888 now includes eight species.

467:
A revision of Othiini. XIX. Two new species of Othius, the first record of the genus from Vietnam, and additional records (Coleoptera: Staphylinidae: Staphylininae). – Othius bihamatus nov.sp. (Pakistan: North-West Frontier Province), a close relative of O. sinuosus Assing, 1998, and O. auster nov.sp. (North Vietnam), the first representative of the genus from the Oriental region and the southernmost record of the genus known at present, are described and illustrated. Additional records of 21 species of Othius Stephens, 1829 and two of Atrecus Jacquelin du Val, 1856 are reported, among them the first record of Atrecus brevicornis Smetana, 1967 since the original description. The distribution of Othius in the East Palaearctic region east of the Himalaya and the individual distributions of eight species are mapped. Othius and the Othiini now include a total of 128 and 145 species and subspecies, respectively.

468:
A new species of Algon from Vietnam (Coleoptera: Staphylinidae: Staphylininae). – Algon fansipanicus nov.sp. (North Vietnam: environs of Sa Pa), a micropterous species of the A. kaiserianus group, is described and illustrated. The A. kaiserianus group is recorded from Vietnam for the first time.

469:
On the Lomechusini fauna of Vietnam (Coleoptera: Staphylinidae: Aleocharinae). – Five species of Lomechusini from the environs of Sa Pa, North Vietnam, are described and illustrated: Amaurodera projecta nov.sp., Drusilla perforans nov.sp., Orphnebius fansipanicus nov.sp., Peltodonia atripalpis nov.sp., and Zyras (Zyras) fansipanicus nov.sp. Three probably undescribed species remain unnamed for want of males. Type material of nine described species is revised and illustrated. The previously unknown male sexual characters of Peltodonia chinensis (Pace, 1998) and Tetrabothrus bicolor Cameron, 1939 are figured for the first time. Two synonymies and two new combinations are proposed: Tetrabothrus japonicus Nakane, 1991 = T. vietnamiculus Pace, 2013; nov.syn.; Zyras neoparageminus Hlaváč, Newton & Maruyama, 2011 = Z. subgeminus Pace, 2012, nov.syn.; Zyras (Zyras) funestus (Dvořak, 1996), nov.comb. (ex Orphnebius Motschulsky, 1858); Zyras (Glossacantha) laocaiensis Pace, 2008, nov.comb. (ex Pella Stephens, 1833). A lectotype is designated for Tetrabothrus bicolor Cameron, 1939. Three species and three genera are recorded from Vietnam for the first time. The Lomechusini fauna of Vietnam is currently represented by 29 species, partly of doubtful status, in seven genera. A checklist of the Lomechusini of Vietnam is provided.

470:
A revision of Zyras Stephens sensu strictu of China, Taiwan, and Hong Kong, with records and (re-)descriptions of some species from other regions (Coleoptera: Staphylinidae: Aleocharinae: Lomechusini). – The species of the nominal subgenus of the lomechusine genus Zyras Stephens, 1835 of China, Taiwan, and Hong Kong are revised. In all, 50 named species, two of them of doubtful identity, are recognized, (re-)described, and illustrated, among them 33 new taxa: Zyras (Zyras) athetoides n. sp. (Sichuan), Z. (Z.) atrapicalis n. sp. (Yunnan), Z. (Z.) atronitens n. sp. (Tibet), Z. (Z.) bangmaicus n. sp. (Yunnan), Z. (Z.) bicoloricollis n. sp. (Yunnan), Z. (Z.) bisinuatus n. sp. (Yunnan), Z. (Z.) caloderoides n. sp. (Yunnan), Z. (Z.) dabanicus n. sp. (Qinghai), Z. (Z.) discolor n. sp. (Fujian), Z. (Z.) exspoliatus n. sp. (Guangxi), Z. (Z.) extensus n. sp. (Yunnan), Z. (Z.) firmicornis n. sp. (Fujian), Z. (Z.) flexus n. sp. (Fujian), Z. (Z.) formosanus n. sp. (Taiwan), Z. (Z.) gilvipalpis n. sp. (Yunnan), Z. (Z.) granapicalis n. sp. (Sichuan), Z. (Z.) hebes n. sp. (Taiwan), Z. (Z.) inexcisus n. sp. (China: Gansu, Qinghai; Russia: Far East, East Siberia), Z. (Z.) lativentris n. sp. (Yunnan), Z. (Z.) maculicollis n. sp. (Yunnan), Z. (Z.) nigrapicalis n. sp. (Yunnan, Sichuan, Jiangxi, Taiwan), Z. (Z.) nigricornis n. sp. (Hubei, Gansu, Shaanxi, Sichuan, Qinghai), Z. (Z.) nigronitens n. sp. (Yunnan), Z. (Z.) pulcher n. sp. (Gansu, Sichuan), Z. (Z.) rectus n. sp. (Yunnan), Z. (Z.) rufapicalis n. sp. (Taiwan), Z. (Z.) rufoterminalis n. sp. (Hubei, Sichuan), Z. (Z.) schuelkei n. sp. (Fujian, Sichuan), Z. (Z.) subobsoletus n. sp. (Sichuan), Z. (Z.) tenebricosus n. sp. (Sichuan, Tibet), Z. (Z.) tenuicornis n. sp. (Taiwan), Z. (Z.) tumidicornis n. sp. (Taiwan), Z. (Z.) volans n. sp. (Taiwan). Previous records of five species from China are considered doubtful. As many as 19 species remain unnamed for want of mature males. A catalogue and a key to species are provided. The distributions of 46 species are mapped. Additional records and (re-)descriptions of 15 species from other – Palaearctic and Oriental – regions are provided. Two species are described for the first time: Z. (Z.) iniquus n. sp. (Pakistan) and Z. (Z.) articollis n. sp. (Laos). Seven synonymies and a new combination are proposed: Zyras beijingensis Pace, 1993 = Z. restitutus Pace, 1993, n. syn.; Zyras alboantennatus Pace, 1986 = Z. sichuanorum Pace, 2012, n. syn.; Z. birmanus Scheerpeltz, 1965 = Z. pseudobirmanus Scheerpeltz, 1965, n. syn.; Z. hongkongensis Pace, 1999 = Z. benenensis Pace, 2001, n. syn.; Z. kambaitiensis Scheerpeltz, 1965 = Z. ferrugineiventris Scheerpeltz, 1965, n. syn., = semiasperatus Scheerpeltz, 1965, n. syn.; Z. wei Pace, 1993 = Z. qingchengensis Pace, 2012, n. syn.; Zyras brignolii (Pace, 1986), n. comb. (ex Drusilla). Lectotypes are designated for Zyras chinkiangensis Bernhauer, 1939 and Z. seminigerrimus Bernhauer, 1933.

471:
On some Lomechusini of the Palaearctic and Oriental regions (Coleoptera: Staphylinidae: Aleocharinae). – Types and additional material of twelve genera, one of them new, of Lomechusini from the Palaearctic and Oriental regions are revised. In all, 70 species are (re-)described and/or illustrated, 51 of them new: Orphnebius bakeri Bernhauer, 1929; O. breviceps Cameron, 1946; O. opticus Cameron, 1946; O. bicuspis sp. n. (Laos; India: Arunachal Pradesh); O. biformis sp. n. (Indonesia: Sumatra); O. cernens sp. n. (Laos); O. dilatatus sp. n. (Laos); O. effeminatus sp. n. (Malaysia); O. extensus sp. n. (Laos); O. fodens sp. n. (Malaysia: Sabah); O. fuscapicalis sp. n. (Laos); O. fusicollis sp. n. (Laos); O. grandicollis sp. n. (Laos); O. integer sp. n. (Laos); O. latitibialis sp. n. (Laos); O. lunatus sp. n. (Laos); O. nigrapicalis sp. n. (Laos); O. reductus sp. n. (Laos); O. retunsus sp. n. (Laos); O. serratus sp. n. (Laos); O. spinans sp. n. (India: Arunachal Pradesh); Orphnebius (Deroleptus) falagrioides Bernhauer, 1929; O. (D.) laticeps Cameron, 1925; O. (D.) multimpressus Assing, 2015; O. (D.) niger (Cameron, 1939); O. (D.) siamensis Cameron, 1939; O. (D.) baccillatus sp. n. (Laos); O. (D.) biimpressus sp. n. (Indonesia: Sumatra); O. (D.) carinatus sp. n. (Laos); O. (D.) cultellatus sp. n. (Thailand, Laos); O. (D.) discrepans sp. n. (China: Yunnan); O. (D.) dispar sp. n. (India: Arunachal Pradesh); O. (D.) gracilior sp. n. (India: Arunachal Pradesh); O. (D.) septemcuspis sp. n. (Laos); O. (D.) sexcarinatus sp. n. (Indonesia: Sumatra); O. (D.) spoliatus sp. n. (Laos); O. (D.) tortus sp. n. (India: Meghalaya); O. (D.) triapicalis sp. n. (China: Sichuan); O. (D.) ulcerosus sp. n. (Malaysia: Sabah); O. (D.) vates sp. n. (Laos); Drusilla bifida sp. n. (Thailand); Rabdotodrusilla pectinata sp. n. (Thailand); Amaurodera meorum Pace, 1992; A. thailandensis Pace, 1986; A. arunica sp. n. (Nepal); A. dentata sp. n. (Thailand); A. disparicollis sp. n. (Indonesia: Sumatra); A. fasciata sp. n. (Thailand); A. gilvios sp. n. (Thailand); A. parvoculata sp. n. (Thailand); A. reticulata sp. n. (Thailand); A. spinans sp. n. (Indonesia: Sumatra); A. varicollis sp. n. (Indonesia: Sumatra); Tetrabothrus borneensis Cameron, 1943; T. indicus Cameron, 1939; T. inflexus Assing, 2015; T. neoguineensis Pace, 2012; T. pubescens Bernhauer, 1915; T. breviatus sp. n. (China: Sichuan); T. collucatus sp. n. (Laos); T. nilgiricus sp. n. (India: Tamil Nadu); T. punctiventris sp. n. (Malaysia: Sabah); T. sulawesicus sp. n. (Indonesia: Sulawesi Utara); Zyras (Zyras) gibbus Pace, 2010; Z. (Z.) illecebrosus Last, 1982; Z. (Z.) quasar Dvořak, 1996; Z. (Z.) porrectus sp. n. (China: Sichuan); Z. (Z.) wunderlei sp. n. (Indonesia: Bali); Pedinopleurus notabilis (Silvestri, 1946); Aenictoides gen. n. derivata sp. n. (Thailand). The intrageneric affiliations of Orphnebius Motschulsky, 1858 and the subgeneric concept currently in use are discussed, and new species (sub-)groups are proposed. The following synonymies are proposed: Deroleptus Bernhauer, 1915 = Megalocephalobius Bernhauer, 1929, syn. n.; Orphnebius breviceps Cameron, 1946 = O. vorax Pace, 2000, syn. n.; Orphnebius niger (Cameron, 1939) = O. turensis Pace, 2012, syn. n.; Drusilla canaliculata (Fabricius, 1787) = Myrmedonia polyporina Gistel, 1857, syn. n.; Drusilla erichsoni (Peyron, 1857) = Myrmedonia aptera Peyron, 1858, syn. n.; Drusilla zyrasoides Dvořak, 1988 = D. truncatella Pace, 2004, syn. n.; Amaurodera kraepelini Fauvel, 1905 = A. rougemonti Pace, 1987, syn. n., = A. smetanai Pace, 1992, syn. n., = A. yaoana Pace, 1992, syn. n.; Tetrabothrus bicolor Cameron, 1939 = T. chinensis Pace, 2012, syn. n.; Tetrabothrus puetzi Assing, 2009 = T. taiwanensis Pace, 2010, syn. n.; Tetrabothrus clavatus Bernhauer, 1915 = T. quadricollis Cameron, 1950, syn. n., = T. japonicus Nakane, 1991, syn. n., = T. vietnamiculus Pace, 2013, syn. n., = T. rubricollis Assing, 2015, syn. n.; Zyras fratrumkadooriorum Pace, 1998 = Z. chumphonensis Pace, 2004, syn. n.; Pedinopleurus notabilis (Silvestri, 1946) = P. setosicauda Dvořák, 1996, syn. n. Lectotypes are designated for Deroleptus niger Cameron, 1939, Orphnebius falagrioides Bernhauer, 1929, Myrmedonia aptera Peyron, 1858, and Tetrabothrus indicus Cameron, 1939. Orphnebius krypticola Pace, 2007 is designated as the correct original spelling. Updated catalogues of Amaurodera Fauvel, 1905 and Tetrabothrus Bernhauer, 1915, as well as keys to the Orphnebius species of Laos and to the Amaurodera species of Thailand are provided. Additional records of 41 named and several unnamed species are reported, among them numerous new country and province records. The distributions of Amaurodera kraepelini and Tetrabothrus clavatus are mapped.

472:
Two new species and additional records of Domene from China and Vietnam (Coleoptera: Staphylinidae: Paederinae). – Two species of Domene Fauvel, 1873 from China are described and illustrated: Domene (Macromene) contiger sp. n. (Guangxi) and D. (M.) exicta sp. n. (Hubei). Additional records of three species are reported from China (Hebei, Hubei, Shaanxi, Sichuan) and North Vietnam. Including the new species, Domene now includes 67 named species, 16 of which have been recorded from China, two from Taiwan, and two from Vietnam.

473:
Six new species, a new name, and additional records of Lathrobium from the Palaearctic region (Coleoptera: Staphylinidae: Paederinae). – Six species of Lathrobium Gravenhorst, 1802 are described and illustrated: Lathrobium kociani nov.sp. (Kazakhstan), L. retunsum nov.sp. (East Nepal) of the L. nepalense group, L. unguiferum nov.sp. (Nepal: Dhaulagiri) of the L. emodense group, L. spinans nov.sp. (Nepal: Manaslu) of the L. pectinatum group, L. jiajinum nov.sp. (China: Sichuan) of the L. bibaculatum group, and L. wolongicum nov.sp. (China: Sichuan) of the L. bibaculatum group. The previously unknown female sexual characters of L. makaluicum Assing, 2013 are illustrated. The preoccupied name Lathrobium abruptum Assing, 2015 is replaced with the nomen novum L. novabruptum. Additional records of 21 species are reported, particularly from Nepal and China. The genus is now represented in the Palaearctic region by 579 species and nine subspecies. The Himalayan fauna currently includes 73 species, all of them micropterous and locally endemic. As many as 211 named species have been recorded from China.

474:
On Oxypoda besucheti Focarile (Coleoptera: Staphylinidae: Aleocharinae: Oxypodini). – Based on a study of type material and numerous additional specimens from several localities in southern Switzerland (Ticino, Grisons) and northern Italy (Lombardia: Alpi Orobie) a new combination and a new synonymy are proposed Tectusa besucheti (Focarile, 1982), nov.comb. (ex Oxypoda Mannerheim, 1830) = T. orobiana Assing, 2012, nov.syn. are proposed. The intraspecific variation of the aedeagal morphology is described, discussed, and illustrated. The distribution of T. besucheti is mapped. The habitat of this alpine species is described and illustrated.

475:
A revision of Geostiba of the West Palaearctic region. XXII. Two new species from Jordan and the Caucaus, and additional records (Coleoptera: Staphylinidae: Aleocharinae). – Two species of Geostiba Thomson, 1858 from the West Caucasus and Jordan are described and illustrated: Geostiba (Sibiota) articarinata nov.sp. (Russia: Krasnodarskiy Kray) and G. (Sipalotricha) jordanica nov.sp., the first representative of the genus to be recorded from Jordan. Additional records of 14 species from the West Palaearctic region are reported.

476:
A new species of Tetrasticta from Laos (Coleoptera: Staphylinidae: Aleocharinae). – Tetrasticta laotica nov.sp. from Laos is described, illustrated, and distinguished from its congeners.

477:
On the Staphylinidae of the Greek island Karpathos (Insecta: Coleoptera). – A study of 931 specimens of Staphylinidae collected in the Greek island Karpathos in December 2015 and January 2016 yielded 59 species, among them eight undescribed island-endemic species and three first records from Greece. A checklist including also previously recorded species is provided. The currently known fauna of Karpathos is composed of 69 species of Staphylinidae and shows stronger affinities to the fauna of Turkey and the Aegean islands off the Turkish coast than to Crete and mainland Greece (including the Pelopónnisos). Ten species (14.5 %) are island endemics, three of which (two of the Pselaphinae and one of the Scydmaeninae) are undescribed. Two myrmecophilous species that were previously considered island endemics of Crete and Rhodos, respectively, are present also in Karpathos. Oxypoda (Atlantoxypoda) bicornuta Assing nov.sp. of the Aleocharinae, Medon carpathicus Assing nov.sp. of the M. petrochilosi subgroup (Paederinae), as well as Cephennium (Phennecium) kerpense Meybohm nov.sp., Euconnus (Tetramelus) kerpensis Meybohm nov.sp., and Stenichnus (Stenichnus) kerpensis Meybohm nov.sp. of the Scydmaeninae are described and illustrated. In an appendix, Oxypoda (Atlantoxypoda) bistirpata Assing nov.sp. (southern Anatolia), a close relative of O. bicornuta, is described and illustrated.

478:
On the Carabidae of the Greek island Karpathos (Insecta: Coleoptera). – A study of 176 specimens of Carabidae collected in the Greek island Karpathos in December 2015 and January 2016 yielded 17 species, among them a new island-endemic species and three first records from Karpathos. The currently known fauna of Karpathos is composed of 48, that of the Karpathos archipelago of 52 species of Carabidae. Three species are island endemics; the status of one taxon is uncertain.

479:
On the Staphylinidae of Turkey XI. Two new species, new synonymies, and additional records (Insecta: Coleoptera). – Two species of Staphylinidae from Turkey are described and illustrated: Liogluta nigrobusta nov.sp. (Artvin) of the Aleocharinae and Vulda substricta nov.sp. (Sinop) of the Xantholinini (Staphylininae). Two synonymies are proposed: Xenochara Mulsant & Rey, 1874 = Rheochara Mulsant & Rey, 1875, nov.syn.; Xantholinus luteipennis Coiffait, 1970 = X. khachikovi Anlaş, 2014, nov.syn. Additional records of 19 named and one unnamed species are reported, several of them considerably expanding the previously known ranges and seven of them representing first records from Turkey.

480:
On Stilicoderus and Stiliderus V. Two new species, additional records, and a catalogue (Coleoptera: Staphylinidae: Paederinae). – Stilicoderus confusus nov.sp. (China: Yunnan; India: Meghalaya) and S. brachypterus nov.sp. (East Nepal), two species that had previously been confounded with S. granulifrons (Rougemont, 1985), are described and illustrated. The distributions of the species of the S. granulifrons group are mapped. Additional records of eleven species of Stilicoderus Sharp, 1889 and four of Stiliderus Motschulsky, 1858 are reported, among them new records from Thailand and Vietnam. A comprehensive catalogue of the 111 species of Stilicoderus and 52 species of Stiliderus currently known is provided.

481:
A revision of Nazeris VIII. Five new species from China and additional records (Coleoptera: Staphylinidae: Paederinae). – Five species of Nazeris Fauvel, 1875 from China are described and illustrated: Nazeris bihamatus nov.sp. (SW-Guangxi), N. latilobatus nov.sp. (N-Guangxi: Maoer Shan), N. obtortus nov.sp. (N-Guangxi: Huaping), N. clavator nov.sp. (Guangdong), and N. latibasalis nov.sp. (Sichuan). Nazeris clavator represents the first record of the genus from Guangdong province. Additional records of 23 described and one undescribed species are reported from the Caucasus, the Himalaya, Vietnam, and China. The genus now includes a total of 241 described species and seven subspecies. It is represented in China by 125 described species.

482:
On some Acanthoglossa and Hypomedon species II. Two new species, a new synonymy, and additional records (Coleoptera: Staphylinidae: Paederinae). – Two species of Hypomedon Mulsant & Rey, 1878 are described and illustrated: H. nasutus nov.sp. (Thailand) and H. bicornutus nov.sp. (Laos). Based on a study of types, the following synonymy is proposed: Acanthoglossa hirta Kraatz; 1859 = Medon sutteri Scheerpeltz, 1957 nov.syn. Records of Hypomedon debilicornis (Wollaston, 1857) and H. galilaeus (Bordoni, 1980) are reported, among them the first records of the H. galilaeus from Pakistan and Taiwan. The enormous range of this species, which now extends from the Middle East to Taiwan, is mapped. Unlike the populations from other regions, the material of H. galilaeus from Taiwan is brachypterous.

483:
On the Staphylinidae of the Greek island Lesbos II, with supplementary notes on the fauna of Samos and Chios (Coleoptera: Staphylinidae). – A field trip to the Greek island Lesbos conducted in spring 2016 yielded a total of 1640 specimens belonging to 169 species of Staphylinidae. Nine new species are described and illustrated: Myllaena lesbia Assing sp.n., Aloconota (Aloconota) aegaea Assing sp.n. (also recorded from Samos), and A. (A.) lesbia Assing sp.n. of the Aleocharinae, Sunius potti Assing sp.n. of the Paederinae, Bryaxis lesbius Brachat sp.n., Bythinus simplicipalpis Brachat sp.n., Protamaurops assingi Brachat sp n., and Tychus lesbius Brachat sp.n. of the Pselaphinae, and Stenichnus (Stenichnus) lesbius Meybohm sp.n. of the Scydmaeninae. The new species of Paederinae, Pselaphinae, and Scydmaeninae are most likely endemic to Lesbos. A comprehensive list of the named and unnamed species currently known from Lesbos is provided. Numerous described species are reported from the island for the first time, eight of them represent first records from Greece. The known staphylinid fauna of Lesbos currently includes 199 species, 184 named and 15 unnamed, unidentified, or tentatively identified. Nine (5 %) of the named species and at least two of the unnamed Pselaphinae and Scydmaeninae are hypothesized to be endemic to the island. Thus, despite greater overall diversity and a generally similar composition, the fauna of Lesbos includes fewer island endemics than that of the geographically close and significantly smaller island Samos. As was to be expected, the fauna of Lesbos displays close affiliations with that of the adjacent Turkish mainland. Several species reach their westernmost distribution limit in this island (some of them also in Chios and Samos). As many as six myrmecophilous species have been recorded from Lesbos, five of them associated with Messor spp. and one with Cataglyphis nodus (Brullé, 1833). In an appendix, Ocalea brachyptera Fagel, 1971 is reported from Chios, the first record of this species from Greece. Aloconota (Aloconota) samia Assing sp.n. (Samos) of the Aleocharinae and Stenichnus (Stenichnus) chius Meybohm sp.n. (Chios) of the Scydmaeninae are described and illustrated.

484:
A revision of Geostiba of the West Palaearctic region. XXIII. On the Sibiota species of the Caucasus region exclusive of Turkey (Coleoptera: Staphylinidae: Aleocharinae). – Seven microphthalmous species of the subgenus Sibiota Casey, 1906, genus Geostiba Thomson, 1858, from Georgia and Armenia are described and illustrated: Geostiba (Sibiota) recta nov.sp. (Georgia: Ratcha and Kvemo-Svanetia provinces); G. (S.) largata nov.sp. (Georgia: Imereti and Ratcha); G. (S.) artifistula nov.sp. (Georgia: Ratcha); G. (S.) unituber nov.sp. (North Armenia: Pambaki range); G. (S.) pambakica nov.sp. (North Armenia: Pambaki range); G. (S.) meghruica nov.sp. (South Armenia: Meghru range); G. (S.) unicuneata nov.sp. (South Armenia: environs of Jermuk). The type locality of G. carinicollis (Eppelsheim, 1878) is revised and G. medea Pace, 1996, previously a junior synonym of G. carinicollis, is revalidated. Additional records of G. bituberculata (Eppelsheim, 1878) and G. kobrisensis Pace, 1996 are reported, and previous records of G. kobrisensis are rectified. A key to the 17 Sibiota currently known from the Caucasus region exclusive of Turkey, all of them microphthalmous and micropterous, is provided. Their distributions are mapped.

485:
New species of Oxypoda from Armenia and Georgia (Coleoptera: Staphylinidae: Aleocharinae). – Six species of Oxypoda Mannerheim, 1830 from Armenia and Georgia are described and illustrated: Oxypoda (Oxypoda) flexa nov.sp. (Armenia), O. (O.) articollis nov.sp. (Georgia), O. (Deropoda) levipunctata nov.sp. (Armenia), O. (Thliboptera) infissoides nov.sp. (Armenia), O. (Bessopora) grandicristata nov.sp. (Armenia), and O. (Sphenoma) subplicata nov.sp. (Armenia). Oxypoda recondita Kraatz, 1856, previously in the subgenus Baeoglena Thomson, 1867, is assigned to Bessopora Thomson, 1859.

486:
On some species of the Quedius obliqueseriatus group, with notes on Q. nivicola (Coleoptera: Staphylinidae: Staphylininae). – New data on several micropterous and locally endemic species distributed in the Caucasus region and allied to Quedius obliqueseriatus Eppelsheim, 1889 are presented. Quedius nigrosuturalis nov.sp. (Northwest Georgia) is described and illustrated. Quedius obliqueseriatus and Q. walteri Korge, 1971, both listed in Microsaurus Dejean, 1833 in the Palaearctic Catalogue, are assigned to the subgenus Raphirus Stephens, 1829. The distributions of Q. boluensis Korge, 1971, Q. smetanai Korge, 1971, and Q. walteri are mapped. Additional records are provided for Quedius nivicola Kiesenwetter, 1858, among them the first records from Turkey.

487:
Revision of the Anaulacaspis species of the Palaearctic region (Coleoptera: Staphylinidae: Aleocharinae). – Types and additional material of the falagriine genus Anaulacaspis Ganglbauer, 1895 are revised. Disregarding one name of doubtful identity, 31 species are recognized in the Palaearctic region, twelve of them new: Anaulacaspis convexa sp. n. (Turkey); A. cristata sp. n. (North Pakistan); A. eminens sp. n. (South Iran, Afghanistan); A. excisa sp. n. (South Iran); A. flavomarginata sp. n. (Albania, Greece); A. gilva sp. n. (North Pakistan); A. iberica sp. n. (Spain); A. pectinata sp. n. (China: Yunnan); A. pseudonigra sp. n. (Turkey); A. reticulata sp. n. (Turkey); A. sinuata sp. n. (Iran); A. truncata sp. n. (Iran). One species, A. inexpectata (Fagel, 1969), is of doubtful generic assignment. The genus and the individual species are (re-)described and illustrated. Twelve synonymies are proposed: Anaulacaspis formosa (Rosenhauer, 1856) = A. cirrosa (Fauvel, 1902), syn. n., = A. algirica (Fagel, 1969), syn. n., = A. elkantarensis (Fagel, 1969), syn. n., = A. macra (Fagel, 1969), syn. n., = A. biskrensis (Fagel, 1969), syn. n., = A. andalusiaca (Fagel, 1969), syn. n.; A. gratilla (Erichson, 1839) = A. lutzi (Reitter, 1909), syn. n.; A. naevula (Erichson, 1839) = A. elegans (Cameron, 1944), syn. n., = A. elegansides Newton, 2015, syn. n.; A. nigra (Gravenhorst, 1802) = A. jonica (Bernhauer, 1910), syn. n., = A. anatolica (Fagel, 1969), syn. n., = A. corcyrana (Fagel, 1969), syn. n. Lectotypes are designated for Aleochara nigra Gravenhorst, 1802, A. picea Gravenhorst, 1802, Falagriola lutzi Reitter, 1909, Falagria laeviuscula Eppelsheim, 1880, F. nigerrima Bernhauer, 1908, and F. beesoni Cameron, 1939. Based on morphological characters, the species are assigned to six species groups. Available data on the zoogeography and natural history of the Anaulacaspis species recorded from the Palaearctic region are compiled. The distributions of the genus as a whole and of the individual species are mapped. A catalogue and a key to species are provided.

488:
On the taxonomy and zoogeography of the species of the Quedius paradisianus group (Coleoptera: Staphylinidae: Staphylininae). – The distributions of four species of the Quedius paradisianus group – Q. paradisianus (Heer, 1839), Q. italicus Gridelli, 1925, Q. collaris Erichson, 1840, and Q. rodopianus Coiffait, 1971 – are revised, mapped, and discussed. While Q. italicus, Q. collaris, and Q. rodopianus have discrete allopatric distributions, that of the widespread and common Q. paradisianus significantly overlaps with the ranges of the other three species. Several new country records are reported. Quedius italicus, previously a subspecies of Q. collaris, is regarded as a distinct species. Lectotypes are designated for Philonthus paradisianus Heer, 1839 and Quedius collaris Erichson, 1840. A key to species and illustrations of relevant diagnostic characters are provided. Data on the natural history are compiled. The vertical distribution and/or the phenology of three species are illustrated.

489:
On the Lomechusini fauna of the East Palaearctic and Oriental regions, with a focus on the genera Orphnebius and Amaurodera (Coleoptera: Staphylinidae: Aleocharinae).. – Types and additional material of 13 genera of Lomechusini from the Palaearctic and Oriental regions are revised. In all, 28 species are (re-)described and/or illustrated, 17 of them new: Orphnebius (Deroleptus) triacuminatus sp. n. (Thailand) of the O. draco group; O. (D.) gracilis sp. n. (South India) and O. (D.) protuberatus sp. n. (South India) of the O. niger group; O. (D.) pertortus sp. n. (India: West Bengal), O. (D.) migrus sp. n. (India: Meghalaya), and O. (D.) reticulipennis sp. n. (Indonesia: Java) of the O. siwalikensis group; Pheidologitonetes bursata sp. n. (South India); Amaurodera bicarinata sp. n. (India: Meghalaya); A. gilvicornis sp. n. (Thailand); A. latisulcata sp. n. (Indonesia: Java); A. brevipes sp. n. (Indonesia: Sumatra); A. migritheca sp. n. (Indonesia: Sumatra); A. longisetosa sp. n. (Malaysia: Sabah); A. calicitheca sp. n. (Malaysia: Sabah); Drusilla lativentris sp. n. (China: Yunnan); Rabdotodrusilla vara sp. n. (India: Meghalaya); Witteia tensa sp. n. (China: Yunnan). The following new combination and synonymy are established: Pheidologitonetes adesi (Pace, 1998), comb. n. (ex Zyrastilbus); Drusilla obliqua (Bernhauer, 1916) = D. palata Assing, 2015, syn. n. Zyrastilbus angkorensis Pace, 2004 is not congeneric with the type species of Zyrastilbus Cameron, 1939; its correct generic assignment (Drusilla?) is currently unknown.  An updated catalogue of the 174 described representatives of Orphnebius Motschulsky, 1858 in the East Palaearctic and Oriental regions is provided; the species are grouped according to a recently proposed intrageneric system. Amaurodera Fauvel, 1905 currently includes 53 described species distributed in the southern East Palaearctic and the Oriental regions. Additional records of 46 named and three unnamed species are reported, among them several new country records. The distributions of two Himalayan Orphnebius species are mapped..

490:
On Zyras sensu strictu in the East Palaearctic and Oriental regions, with a focus on the faunas of the Himalaya, India, Stri Lanka, Thailand, and Sulawesi (Coleoptera: Staphylinidae: Aleocharinae: Lomechusini). – The species of the nominal subgenus of the lomechusine genus Zyras Stephens, 1835 of the Himalaya, India, Sri Lanka, Thailand, and Sulawesi are revised. Additional species from other parts of the East Palaearctic and Oriental regions are addressed. In all, 40 species are described and/or illustrated, 14 of them for the first time: Zyras (Zyras) ambulans spec. nov. (Thailand), Z. (Z.) brevilobatus spec. nov. (Thailand), Z. (Z.) densihirtus spec. nov. (Sulawesi Utara), Z. (Z.) densissimus spec. nov. (Sulawesi Utara), Z. (Z.) latilobatus spec. nov. (South India), Z. (Z.) longilobatus spec. nov. (India: Meghalaya), Z. (Z.) luteipes spec. nov. (India: Meghalaya), Z. (Z.) morulus spec. nov. (Nepal: Dhaulagiri, Annapurna), Z. (Z.) nigrihirtus spec. nov. (Sulawesi Utara), Z. (Z.) parahirtus spec. nov. (Borneo), Z. (Z.) parvicollis spec. nov. (Thailand), Z. (Z.) russiceps spec. nov. (Thailand, Malaysia), Z. (Z.) titan spec. nov. (Sulawesi Utara), and Z. (Z.) truncatus spec. nov. (Nepal: Dhaulagiri). As many as 28 synonymies are proposed: Zyras alternans (Cameron, 1925) = Z. optimus Cameron, 1939, syn. nov.; Z. bartolozzii Pace, 2003 = Z. alboterminalis Pace, 2008, syn. nov.; Z. bettotanus Cameron, 1930 = Z. drescheri Cameron, 1939, syn. nov., = Z. atrapicalis Assing, 2016, syn. nov.; Z. brignolii (Pace, 1986) = Z. thainiger Pace, 2012, syn. nov.; Z. castaneus (Motschulsky, 1861) = Z. adulescens (Pace, 1987), syn. nov.,  = Z. britannorum Pace, 1992, syn. nov., = Z. fratrumkadooriorum Pace, 1998, syn. nov., = Z. chumphonensis Pace, 2004, syn. nov., = Z. dibrugarhensis Pace, 2011, syn. nov.; Z. preangeranus Cameron, 1939 = Z. louwerensi Cameron, 1939, syn. nov., = Z. chinkiangensis Bernhauer, 1939, syn. nov., = Z. setosipennis Scheerpeltz, 1965, syn. nov., = Z. alboantennatus Pace, 1986, syn. nov., = Z. sichuanorum Pace, 2012, syn. nov.; Z. geminus (Kraatz, 1859) = Z. indicus Cameron, 1944, syn. nov., = Z. shiva Pace, 1992, syn. nov., = Z. manjushri Pace, 1992, syn. nov., = Z. hongkongensis Pace, 1999, syn. nov., = Z. benenensis Pace, 2001, syn. nov., = Z. parageminus Pace, 2010, syn. nov., = Z. neoparageminus Hlaváč, Newton & Maruyama, 2011, syn. nov., = Z. subgeminus Pace, 2012, syn. nov., = Z. articollis Assing, 2016, syn. nov.; Z. parageminus Pace, 1998 = Z. nameriensis Pace, 2011, syn. nov.; Z. pindarae (Champion, 1921) = Z. ruficauda Cameron, 1939, syn. nov.; Z. proximus Cameron, 1939 = Z. drugmandi Pace, 2004; syn. nov. Zyras novinversus nom. n. is proposed for the preoccupied name Z. inversus Pace, 2012, syn. nov. Lectotypes are designated for Zyras exasperatus Schubert, 1908, Z. drescheri Cameron, 1939, Z. gratellus Cameron, 1939, Myrmedonia perforata Champion, 1921, and Hygroptera castanea Motschulsky, 1861. Remarkable cases of colour polymorphism and of sexual dimorphism are discussed. A key to the Zyras sensu strictu species of the Himalaya, India, and Sri Lanka, a key to the species recorded from Thailand, and an updated catalogue of the Zyras sensu strictu species of the Palaearctic and Oriental regions are provided. Additional records of numerous named and several unnamed species are reported. The revised distributions of 37 species are mapped. The subgenus is currently represented in the Palaearctic and Oriental regions by a total of 124 described species. The countries with the greatest diversity are China (46 named species), Malaysia (20), India (20), Indonesia (18), Thailand (13), and Nepal (12). The subgenus is reported from Sulawesi for the first time (five named and two unnamed species). Based on a revision of type material, eleven species are not included in, or excluded from, Zyras sensu strictu, and Zyras unicolor Cameron, 1939 is tentatively moved to Drusilla Leach, 1819..

491:
On the Staphylinidae of the Greek island Kos, with an appendix on Carabidae and additional records from other islands (Insecta: Coleoptera). – A study of 567 specimens of Staphylinidae collected in the Greek island Kos in April 2012 and December 2016 yielded 54 species, among them five first records from Greece. One of them is Quedius curtidens Smetana, 1967, a species previously known only from Adana province (Turkey) and also reported from Lebanon, Syria, and Israel for the first time; its distribution is mapped. A checklist of the species recorded from Kos is provided. The known staphylinid fauna is represented by 58 species and shows strong affinities to the fauna of the close Turkish mainland. It is primarily composed of widespread West Palaearctic (40 %), East Mediterranean (36 %), South Anatolian (11 %), or widespread Mediterranean (8 %) species. In an appendix, records of 15 species are reported from Cyprus, Rhodos, Samos, and Lesbos. Moreover, 29 species of Carabidae (177 specimens) are reported from Kos; eight of these species are recorded from the Aegean Islands for the first time.

492:
On Quedius coloratus Fauvel, 1875 and allied species, with an appendix on Quedius species collected in Greece iwth subterranean pitfall traps, and a new synonymy (Coleoptera: Staphylinidae: Staphylininae). – A revision of material previously identified as Quedius coloratus Fauvel, 1875 revealed that it is composed of at least five species: Q. (Raphirus) coloratus (Middle East); Q. (R.) hellenicus nov.sp. (Greece); Q. (R.) hebes nov.sp. (South Anatolia); Q. (R.) spiculatus nov.sp. (North Anatolia); Q. (R.) carpathius nov.sp. (Karpathos). These species, as well as one male possibly representing a sixth species (Greece: Thessalía) are described and illustrated. Available evidence suggests that their habitats are subterranean and their distributions are allo- or parapatric. All the named species are presumably capable of flight. A key to species is provided. In an appendix, additional records of three Quedius species collected in Greece with subterranean pitfall traps are reported. One of these species is reported from Greece for the first time. The distributions of the species allied to Q. coloratus, of Q. (Microsaurus) bernhaueri Rambousek, 1915, and of Q. (Raphirus) endogeus Assing, 2007 are mapped. Quedius endogeus, a species previously known only from two localities, is widespread in Greece and evidently a true inhabitant of the "Superficial Subterranean Habitat" (MSS stratum). Quedius crnagoricus Coiffait, 1980, nov.syn., is placed in synonymy with Q. paradisianus (Heer, 1839).

493:
A revision of the Alevonota species of the Palaearctic region. II. A new species from Cyprus and additional records (Coleoptera: Staphylinidae: Aleocharinae). – Alevonota flexa nov.sp. (Cyprus) is described and illustrated. Additional records of four species are reported, among them several new country records. Alevonota japonica (Cameron, 1933), a species with a remarkable sexual dimorphism of head shape and eye size, is illustrated, based on new material from Hokkaido.

494:
Taxonomic and faunistic notes on some West Palaearctic and Middle Asian Xantholinini, with a revalidation and new synonymies (Coleoptera: Staphylinidae: Staphylininae). – Xantholinus motschulskyi Bordoni, 1999, previously a synonym of X. reitteri Coiffait, 1966, is revalidated. The internal structures of the aedeagus of X. motschulskyi, X. reitteri, and X. corallinus Reitter, 1901 are illustrated. A lectotype is designated for X. corallinus. Six synonymies are proposed: Gyrohypnus punctulatus (Paykull, 1789) = Gyroyhypnus quadratus Stephens, 1833, nov.syn.; Stenistoderus cephalotes cephalotes (Kraatz, 1858) = S. cephalotes armeniacus (Coiffait, 1966), nov.syn.; Xantholinus linearis (Olivier, 1795) = X. styriacus Grimmer, 1841, nov.syn.; X. gridellii Coiffait, 1956 = X. tronqueti Bordoni, 2016, nov.syn.; X. khnzoriani Coiffait, 1966 = X. caucasicus Bordoni, 1975, nov.syn., = X. iablokoffi Coiffait, 1975, nov.syn. A neotype is designated for Gyroyhypnus quadratus Stephens, 1833. New records of 37 species of Xantholinini are reported. The revised distributions of X. reitteri and X. motschulskyi are mapped.

495:
Two new species and additional records of micropterous Paederus from China and Taiwan (Coleoptera: Staphylinidae: Staphylininae). – Two species of Paederus Fabricius, 1775 from China and Taiwan are described and illustrated: Paederus (Gnathopaederus) furcillatus nov.sp. (China: Guizhou) and P. (Megalopaederus) nigricrus nov.sp. (Taiwan: Taitung Hsien). Additional records of eight species of Harpopaederus Scheerpeltz, 1957 (two species), Gnathopaederus Chapin, 1927 (one species), Megalopaederus Scheerpeltz, 1957 (three species), and of the P. biacutus group (two species) are reported. The genus is currently represented in China and Taiwan by 51 and 25 species respectively.

496:
A revision of Leptobium Casey. VIII. A new species from Iran, a new synonymy, and additional records (Coleoptera: Staphylinidae: Paederinae). – Leptobium iranicum nov.sp. (Iran: Yazd province) is described and illustrated. One synonymy is proposed: Leptobium carinatum Assing, 2005 = nabozhenkoi Anlaş, 2012, nov.syn. Additional records of 25 species and subspecies are reported. The genus now includes a total of 67 species and two subspecies.

497:
Four new species and additional records of Lathrobium from Nepal and China (Coleoptera: Staphylinidae: Paederinae). – Four species of Lathrobium Gravenhorst, 1802 are described and illustrated: Lathrobium kongmaicum nov.sp. (East Nepal: Kongma Danda), L. latilobatum nov.sp. (East Nepal: Kongma Danda), and L. paulitortum nov.sp. (East Nepal: Solu Khumbu) of the L. nepalense group, and L. piraticum nov.sp. (China: Sichuan: Wolong) of the L. fissispinosum group. Additional records of three species from China and three species from Nepal are reported, among them the first recent records of L. nepalorientis Coiffait, 1984 and L. khumbuense Coiffait, 1982. The genus is now represented in the Palaearctic region by 595 species and nine subspecies. The Himalayan fauna currently includes 76 described species, all of them micropterous and locally endemic. As many as 220 named species have been recorded from China.

498:
A revision of Palaearctic Lobrathium. VI. Two new species from Armenia and Taiwan, and additional records (Coleoptera: Staphylinidae: Paederinae). – Two species of Lobrathium Mulsant & Rey, 1878 are described and illustrated: L. ancoriferum nov.sp. (Armenia) and L. biuncinatum nov.sp. (Taiwan). Additional records of 13 species are presented. Lobrathium is now represented in the Palaearctic region (including Myanmar and Vietnam) by 144 described species and one subspecies.

499:
A revision of Palaearctic Nazeris IX. Three new species from Nepal and additional records (Coleoptera: Staphylinidae: Paederinae). – Three species of Nazeris Fauvel, 1875 of the N. flavocaudatus group from East Nepal are described and illustrated: Nazeris biapicalis nov.sp., N. hebes nov.sp., and N. trapezilobus nov.sp. Additional records of three previously described species are reported from East Nepal and Lebanon. The pronounced intraspecific variation of external and sexual characters of N. ammonita (Saulcy, 1865) is discussed and illustrated. The genus now includes a total of 250 described species and seven subspecies. It is represented in the Himalaya (exclusive of China) by 52 described species.

500:
A revision of Neosclerus Cameron VI. Thre new species from India, Thailand, and Vietnam (Coleoptera: Staphylinidae: Paederinae). – Three species of Neosclerus Cameron, 1924 are described and illustrated: N. anguliceps nov.sp. (South India), a species of uncertain affiliations, N. concavus nov.sp. (Thailand) of the N. brevispinosus group, and N. vietnamensis (Vietnam) of the N. brevispinosus group. The latter species represents the second record of the genus from Vietnam. Including the new species, the genus now includes 47 species distributed in the southern East Palaearctic and Oriental regions.

501:
The first species of Chinecallicerus from Sichuan, China (Coleoptera: Aleocharinae: Geostibini). – Chinecallicerus pinnatus nov.sp. (China: Sichuan: Wolong), the first representative of the genus to be recorded from outside Yunnan, is described, illustrated, and distinguished from its congeners. An updated key to the species of Chinecallicerus Assing, 2004 is provided. The genus currently includes seven species, six of which are representely solely by their respective holotypes.

502:
A revision of types and additional material of Staphylinidae from the Iablokoff-Khnzorian collection (Insecta: Coleoptera). – Types and additional material from the Iablokoff-Khnzorian collection are revised. The following synonymies are proposed: Olophrum puncticolle Eppelsheim, 1880 = O. erevanicum Iablokoff-Khnzorian, 1964, nov.syn.; Astenus procerus (Gravenhorst, 1806) = Astenus silvicola Iablokoff-Khnzorian, 1961, nov.syn.; Myrmoecia reitteri (Eppelsheim, 1881) = Zyras argus Iablokoff-Khnzorian, 1960, nov.syn.; Aleochara (Xenochara) rambouseki Likovský, 1964 = A. addenda Likovský, 1981, nov.syn., = A. hamulata Assing, 2009, nov.syn. The holotypes of Rugilus armeniacus (Coiffait, 1970), Aleochara polychroma Iablokoff-Khnzorian, 1966, and A. nobilis Likovský, 1965 are illustrated. The latter is moved to the subgenus Xenochara Mulsant & Rey, 1874 (ex Heterochara Mulsant & Rey, 1874). Seven species of Staphylinidae are reported from Armenia and one from Slovenia for the first time. The record of one species from Armenia is based on a misentification.

503:
On some Athetini from Armenia and adjacent regions (Coleoptera: Aleocharinae). – Nine species of Athetini from Armenia are described and illustrated, eight of them for the first time: Atheta (Atheta) brevapicalis nov.sp. (South Armenia); A. (A.) hamulata nov.sp. (South Armenia); A. (Dimetrota) bispinosa nov.sp. (Armenia, Georgia); A. (D.) senticollis nov.sp. (North Armenia); A. (Oreostiba) altiviva Benick, 1974 (widespread in Armenia); A. (O.) semialba nov.sp. (widespread in Armenia); A. (O.) abscisa nov.sp. (widespread in Armenia); A. (O.) brevitheca nov.sp. (widespread in Armenia); Liogluta armeniaca nov.sp. (Central and South Armenia). Three synonymies are proposed: Atheta hansseni Strand, 1943 = A. brachati Assing, 2013, nov.syn.; Liogluta granigera (Kiesenwetter, 1850) = L. trigemina (Eppelsheim, 1880), nov.syn.; L. funesta Eppelsheim, 1890 = L. nigrobusta Assing, 2016, nov.syn. A lectotype is designated for Homalota trigemina Eppelsheim, 1880.

504:
On Stilicoderus VI. Two new species from South Vietnam and additional records (Coleoptera: Paederinae). – Two species of Stilicoderus Sharp, 1889 from South Vietnam are described and illustrated: Stilicoderus bisinuatus nov.sp. of the S. granulifrons group and S. harpago nov.sp. of the S. discalis group. Additional records of five species are reported, among them a new record from Bhutan. The genus now includes a total of 113 species.

506:
A revision of Nazeris X. The first record of the genus from South Vietnam and additional records from Thailand (Coleoptera: Staphylinidae: Paederinae). – Nazeris raptus nov.sp. from South Vietnam is described and illustrated. This record is of special zoogeographic significance, as it is situated nearly 6.5 degrees of latitude farther south than the previous southernmost record of the genus as a whole, thus casting doubt on the previous classification of the distribution of Nazeris Fauvel, 1873 as essentially South Palaearctic. Additional records of two species from Thailand are reported. An updated map illustrating the distribution of the genus as a whole is provided. Including the new species, Nazeris now includes 255 described species and seven subspecies.

507:
Two new species of Gnathymenus from Ecuador (Coleoptera: Staphylinidae: Paederinae). – Two species of the dolicaonine genus Gnathymenus Solier, 1849 from Ecuador are described and illustrated, Gnathymenus arboreus nov.sp. and G. spinifer nov.sp., both of them macropterous and collected by canopy fogging in the same locality in Pastaza province. The genus now includes 77 species, 16 of which are known from Ecuador. Additional records of Gnathymenus apterus Solier, 1849 are reported from Chile.

508:
On the micropterous Quedius (Raphirus) species with a punctate scutellum of Turkey (Coleoptera: Staphylinidae: Staphylininae). – Four micropterous species of Quedius Stephens, 1829, subgenus Raphirus Stephens, 1829, with a punctate scutellum are reported from North Turkey, one of them unidentified. Quedius brevalatus nov.sp. (Armenia; Turkey: Rize) is described and illustrated. Quedius omissus Coiffait, 1977, a species previously known only from the West Caucasus, is reported from Turkey (Rize) for the first time. Three synonymies are proposed: Quedius boops (Gravenhorst, 1802) = Philonthus boops tauricus Nordmann, 1837, nov.syn., = Quedius crius Tottenham, 1948, nov.syn. (previously a synonym of Q. reitteri Gridelli, 1925), = Quedius boops islandicus Fagel, 1960, nov.syn.

509:
A mass dispersal event of Quedius hellenicus (Coleoptera: Staphylinidae: Staphylininae). – In spring 2017, a mass dispersal event of Quedius hellenicus Assing, 2017, the first observation of such a behaviour in the genus, was witnessed in the Greek island Corfu. Additional evidence is presented suggesting that Q. hellenicus inhabits a subterranean habitat and that this most likely also applies to other representatives of the Q. coloratus group.

510:
On the Leptusa fauna of the Caucasus region (Coleoptera: Staphylinidae: Aleocharinae). – Five species of Leptusa Kraatz, 1858 from Georgia and North Iran are described and illustrated: L. (Neopisalia) substricta nov.sp. (Georgia: Shida Kartli); L. (N.) svanetica nov.sp. (Georgia: Svaneti); L. (N.) migrituber nov.sp. (Georgia: Svaneti); L. (N.) triangulata nov.sp. (Georgia: Adjara); L. (N.) longalata nov.sp. (Iran: Mazandaran). The genus is represented in the Caucasus region including Northeast Turkey (from Ordu to the border with Georgia), Georgia, Armenia, Azerbaijan, and the Russian part of the Greater Caucasus by 39 species in five subgenera; 28 species, two of them of doubtful identity, belong to the subgenus Neopisalia Scheerpeltz, 1966. Three zoogeogeographically implausible or revised previous records are rectified. Leptusa kaszabi Pace, 1981 is moved from the subgenus Heteroleptusa Pace, 1989 to Neopisalia. A zoogeographic study of the Leptusa fauna of the Caucasus region revealed that locally endemic species are confined to the moister western half of the region eastwards approximately to 43°30' eastern longitude. Moreover, little is known about the fauna of Abkhazia, which probably hosts an unknown number of unnamed species. A catalogue of the Leptusa species of the Caucasus region and maps illustrating the currently known distributions of 33 species are provided. Additional records of 19 described species are reported, among them several new country records.

511:
On the Geostiba fauna of Armenia (Coleoptera: Staphylinidae: Aleocharinae). – Three microphthalmous species of the subgenus Sibiota Casey, 1906, genus Geostiba Thomson, 1858, from Armenia are described and illustrated: Geostiba (Sibiota) tigrani nov.sp. (North Armenia: mountains to the north of Sevan lake); G. (S.) kalavanica nov.sp. (North Armenia: Kalavan mountain near the northwestern tip of Sevan lake); G. (S.) immutata nov.sp. (South Armenia: Barkushati range). Geostiba (Geostiba) sororcula Assing, 2001, a species previously known only from Northeast Turkey, is reported from Armenia for the first time. Like its close relative, the widespread G. circellaris (Gravenhorst, 1806), this species is wing-dimorphic. At present, the known Geostiba fauna is composed of nine described species in three subgenera; one or two additional species are unnamed (males unavailable). Seven described and the unnamed species, all of them microphthalmous, micropterous, and more or less locally endemic, belong to the subgenus Sibiota; they are subject to remarkable intraspecific variation of the male secondary sexual characters. A key to species and distribution maps are provided.

512:
A revision of Geostiba of the West Palaearctic region. XXV. New species from Georgia and Greece, and additional records (Coleoptera: Staphylinidae: Aleocharinae). – Four species of Geostiba Thomson, 1858 from Georgia and Greece are described and illustrated: Geostiba (Tropogastrosipalia) xirosica nov.sp. (Greece: Evvoia); G. (T.) svanetica nov.sp. (Georgia: Svaneti); G. (T.) gibberiventris nov.sp. (Georgia: Mtskheta-Mtianeti); G. (Sibiota) granisuturalis nov.sp. (Georgia: Svaneti). Additional records of seven species are reported. The distributions of the eight species of the subgenus Tropogastrosipalia Scheerpeltz, 1951 recorded from the Caucasus region, those of the nine species of Sibiota Casey, 1906 known from the West and Central Greater Caucasus, and the distribution of G. (Chondridiosipalia) cingulata (Eppelsheim, 1878) in the Caucasus region are mapped. Supplements to recently published keys to the Geostiba fauna of the West Palaearctic region are provided.

513:
On the Ischnosoma fauna of Georgia (Coleoptera: Staphylinidae: Tachyporinae). – Three species of the Ischnosoma spelaeum group are described and illustrated, all of them most likely with very restricted distributions: Ischnosoma acutum spec. nov. (Georgia: Svaneti, Racha); I. barbigerum spec. nov. (Georgia: Svaneti); I. molle spec. nov. (Georgia: Adjara). Type material of I. major (Luze, 1901) and I. caucasicum Kocian, 1997 is revised. Additional records of previously described species of Ischnosoma Stephens, 1829 are reported from Georgia, partly also from other regions. Seven species of the genus have reliably been recorded from Georgia, two species of the I. pictum group and widespread, five of the I. spelaeum group and with restricted distributions. The presence of an additional species of the I. spelaeum group, I. caucasicum, requires confirmation. The distributions of the species of the I. spelaeum group are revised and mapped. Several previous literature records are zoogeographically implausible and consequently probably erroneous.

514:
On the Zyras sensu strictu in the East Palaearctic and Oriental regions III, with a focus on the Southeast of Continental Asia and the Sunda Islands (Coleoptera: Staphylinidae: Aleocharinae: Lomechusini). – Species of the nominal subgenus of the lomechusine genus Zyras Stephens, 1835 of the southeastern Oriental region are revised. Eighteen species are (re-)described and/or illustrated, six of them for the first time: Zyras (Zyras) latibasalis spec. nov. (Java, tentatively recorded also from Laos), Z. (Z.) rutrilobatus spec. nov. (Laos), Z. (Z.) parvilobatus spec. nov. (Laos), Z. (Z.) hirtiventris spec. nov. (Laos, Vietnam), Z. (Z.) hlavaci spec. nov. (Malaysia: Pahang, Selangor), Z. (Z.) lunatus spec. nov. (Malaysia: Selangor). Eight synonymies are proposed: Zyras bryanti Cameron, 1943 = Z. mortuorum Pace, 1990, syn. nov., = Z. paederinus Pace, 2008, syn. nov.; Zyras nigerrimus Cameron, 1943 = Z. bartolozzii Pace, 2003, syn. nov., = Z. alboterminalis Pace, 2008, syn. nov.; Z. granulipennis Cameron, 1930 = Z. pervariolosus Pace, 2008, syn. nov.; Z. matangensis Cameron, 1943 = Z. daiaccorum Pace, 2008, syn. nov.; Z. montanus (Bernhauer, 1915) = Z. variolatus Pace, 2003, syn. nov.; Z. preangeranus Cameron, 1939 = Z. quadriterminalis Pace, 2008, syn. nov. One species is excluded from Zyras: Myrmedonota modiglianii (Cameron, 1925), comb. nov. Lectotypes are designated for Zyras elegantulus Cameron, 1939, Z. semirufus Cameron, 1939, and Z. flavus Cameron, 1939. Additional records of nine named species are reported. A key to the species recorded from the southeastern Oriental region and an updated catalogue of the species of the Palaearctic and Oriental regions are provided. The subgenus is currently represented in the Palaearctic and Oriental regions by a total of 122 described species. Twenty-two species have been recorded from Sunda Islands. The countries with the greatest diversity are China (46 named species), India (20), Indonesia (19), Malaysia (15), Thailand (14), Nepal (12), Laos (11), and Vietnam (8).

515:
A revision of Sunius XVI. Two new species from Iran and Iraq (Coleoptera: Staphylinidae: Paederinae). – Two species of Sunius Stephens, 1829 are described and illustrated: S. reuteri spec. nov. (North Iraq), the first record of the genus from Iraq, and S. concurvatus spec. nov. (South Iran). Additional records of six previously described species are reported from the West Palaearctic region. Updated distribution maps are provided for S. khnzoriani (Coiffait, 1970) and S. fulgocephalus (Coiffait, 1970). The genus is currently represented in the Palaearctic region by 138 described species and two subspecies.

516:
On the Staphylinidae of Crete III. The first records of endogean fauna (Coleoptera: Staphylinidae: Leptotyphlinae, Aleocharinae). – Endogean Staphylinidae were previously unknown from the Greek island Crete. During a field trip conducted to East Crete in December and January 2017/2018 four species of endogean Staphylinidae, three of the Leptotyphlinae and one of the Aleocharinae, were collected using the soil-washing method. Two of the leptotyphline species are described and illustrated: Kenotyphlus creticus nov.sp. and Allotyphlus (Moreotyphlus) candicus nov.sp. The third species of Leptotyphlinae belongs to the genus Gynotyphlus Coiffait, 1955 and remains undescribed for want of males. Typhlocyptus pandellei Saulcy, 1878 (Aleocharinae) is reported from Crete and Montenegro for the first time.

517:
The third hypogean Domene species from Greece (Coleoptera: Staphylinidae: Paederinae). – TDomene (Domene) vailatii nov.sp. (Greece: Thesprotía), the third hypogean representative of Domene Fauvel, 1873 from Greece, is described and illustrated. The species is apparently subject to remarkably biased sex ratio; only one of the 22 type specimens is a male. The distributions of the four hypogean Domene species known from Greece and Albania are mapped.

518:
A revision of Othiini. XX. Two new species of Othius, the first record of the genus from Myanmar, and additional records (Coleoptera: Staphylinidae: Staphylininae). – Othius arater nov.sp. (China: South Sichuan) and O. burmensis nov.sp. (Northeast Myanmar), the first representative of the genus recorded from Myanmar, are described and illustrated. New illustrations of the external and male sexual characters of O. discrepans Assing, 1999 are provided. Additional records of ten species are reported. The distributions of three Caucasian species are mapped. Othius Stephens, 1829 and the Othiini now include a total of 130 and 147 species and subspecies, respectively.

519:
On Hypnogyra laevissima (Reitter, 1898) (Coleoptera: Staphylinidae: Staphylininae: Xantholinini). – Based on a revision of types and additional material, Hypnogyra laevissima (Reitter, 1898), a name recently placed in synonymy with the widespread H. angularis (Ganglbauer, 1895), is revalidated. The external and male sexual characters are illustrated and compared with those of H. angularis. The known distribution of H. laevissima, which is confined to East Azerbaijan and North Iran, is mapped.

520:
Six new species of Chinecallicerus from China, with a new synonymy (Coleoptera: Staphylinidae: Aleocharinae: Geostibini). – Six species of Chinecallicerus Assing, 2004 are described and illustrated: Chinecallicerus carinatus nov.sp. (China: North Sichuan), C. orbiculatus nov.sp. (China: South Sichuan), C. transversus nov.sp. (China: North Sichuan), C. granulosissimus nov.sp. (China: East Qinghai), C. grandicollis nov.sp. (China: Northwest Yunnan), and C. trituberculatus nov.sp. (China: Northwest Yunnan). The following synonymy is proposed: Chinecallicerus schuelkei Assing, 2004 = Aloconota sinica Pace, 2011, nov.syn. An updated key to the Chinecallicerus species is provided. Including the new species, the genus now includes 13 species, eight of which have been recorded from the Chinese province Yunnan, four from Sichuan, and one from Qinghai. All the species are known only from their respective type localities, nine of them are representely solely by their respective holotypes. The distributions of all the species of the genus are mapped.

521:
On the Nepalota fauna of China II. Two new synonymies and an additional record (Coleoptera: Staphylinidae: Aleocharinae). – Two synonymies are proposed: Nepalota cuneata Assing, 2015 = N. gibbera Pace, 2017, nov.syn.; N. tuberifera Assing, 2015 = N. tengchongensis Pace, 2017, nov.syn. A new record of N. chinensis Pace, 1998 from the Chinese province Jiangxi is provided.

522:
A new Atheta species from the West Caucasus and a redescription of A. brevapicalis (Coleoptera: Staphylinidae: Aleocharinae). – Atheta ("Alaobia") specicola nov.sp. (Russia: Krasnodarskiy Kray), a cavernicolous species closely allied to A. trinotata (Kraatz, 1856), and A. brevapicalis Assing & Vogel, 2017 are (re-)described and illustrated. Newly collected material revealed that the type series of A. brevapicalis is composed of two species. Additional records of A. brevapicalis, including the first records from Iran, are reported; the distribution of the species is mapped.

523:
New species and additional records of Cephalocousya and Drepasiagonusa from China and Russia (Coleoptera: Staphylinidae: Aleocharinae: Oxypodini). – One species of Cephalocousya Lohse, 1971 and four species of Drepasiagonusa Pace, 2012 are described and illustrated: Cephalocousya yunnanica nov.sp. (China: North Yunnan), the first record of the genus from China; Drepasiagonusa procera nov.sp. (China: North Yunnan); D. tibetica nov.sp. (China: North Yunnan); D. angulata nov.sp. (China: Qinghai, Gansu); D. feldmanni nov.sp. (Russia: East Siberia). The previously unknown aedeagus of D. smetanai Pace, 2012, the type species and previously the sole representative of the genus, is illustrated. Two additional Drepasiagonusa species from Shaanxi are represented exclusively by females and remain unnamed. A key to the described species of Drepasiagonusa is provided. The currently known distribution of the genus is mapped. Cephalocousya nivicola is reported from Kyrgyzstan for the first time.

524:
Six new species of Trichoglossina from China (Coleoptera: Staphylinidae: Aleocharinae: Oxypodini). – Six species of Trichoglossina Pace, 1987 from China are described and illustrated: Trichoglossina aptera nov.sp. (North Sichuan), T. alticola nov.sp. (North Sichuan), T. acutissima nov.sp. (North Sichuan), T. volans nov.sp. (North Sichuan), T. discolor nov.sp. (Shaanxi), and T. bifida (Northeast Yunnan). The distributions of these species are mapped. The genus, which has been recorded only from Nepal and China, now includes 39 species.

525:
Three new species of Oxypoda from Spain, Armenia, and Ukraine, with notes on the fauna of Armenia (Coleoptera: Staphylinidae: Aleocharinae). – Three species of Oxypoda Mannerheim, 1830 from Spain, Armenia, and Ukraine are described and illustrated: Oxypoda (Bessopora) parvioculata nov.sp. (Spain: Andalucía: Sierra Nevada), O. (Podoxya) verminata nov.sp. (Armenia), and O. breviata nov.sp. (Ukraine). Additional records of 18 species are reported from Armenia, 13 of them for the first time. The currently known Armenian Oxypoda fauna is composed of at least 22 species, 20 of them named and 6 exclusive. The currently known distributions of six species are mapped.

526:
On the Aleochara subgenera Ceranota and Xenochara. IV. A revision of types, a new species, and additional records (Coleoptera: Staphylinidae: Aleocharinae). – Aleochara (Ceranota) plicelytrata nov.sp. (Armenia, Georgia) of the A. subtumida group is described and illustrated. Aleochara (Xenochara) citellorum Kirshenblat, 1935 and A. (X.) jacobsoni Kirshenblat, 1935 are revised, redescribed, and illustrated; lectotypes are designated for both species. They are assigned to the A. parvicornis group. Additional records are reported for 13 species of Ceranota Stephens, 1839 and 18 species of Xenochara Mulsant & Rey, 1874, among them 17 first records for Armenia (six), Georgia (three), Ukraine (three), Russia (two), Lebanon (two), Greece (one), Iran (one), and Kashmir (one). The distributions of six Ceranota and one Xenochara species are mapped.

527:
On the taxonomy and zoogeography of some West Palaearctic Quedius species, with a focus on the East Mediterranean and the species allied to Quedius umbrinus and Q. nivicola (Coleoptera: Staphylinidae: Staphylininae). – Based on a revision of type material, as well as on studies of original descriptions and intraspecific variation, 21 synonymies are proposed in the subgenus Raphirus Stephens, 1829 of the genus Quedius Stephens, 1829: Quedius umbrinus Erichson, 1839 = Q. cyanescens Mulsant & Rey, 1876, nov.syn., = Q. bulgaricus Scheerpeltz, 1937, nov.syn., = Q. cyprusensis Last, 1955, nov.syn., = Q. freyi Scheerpeltz, 1956, nov.syn., = Q. maronitus Coiffait, 1963, nov.syn., = Q. gueorguievi Coiffait, 1967, nov.syn.; Quedius hermonensis Coiffait, 1963 = Q. coiffaitianus Fagel, 1968, nov.syn., = Q. rugosipennis Fagel, 1969, nov.syn.; Quedius illyricus Wendeler, 1928 = Q. paganettii Bernhauer, 1936, nov.syn., = Q. schipkanus Scheerpeltz, 1937, nov.syn., = Q. pseudopyrenaeus Coiffait, 1967, nov.syn.; Quedius nemoralis Baudi di Selve, 1848 = Q. safaensis Fagel, 1968, nov.syn., = Q. safaensis ormanus Fagel, 1971, nov.syn., = Q. nemoralis erinci Korge, 1971, nov.syn.; Quedius limbatus (Heer, 1839) = Q. scheerpeltzianus Fagel, 1968, nov.syn.; Quedius suturalis Kiesenwetter, 1845 = Q. humeralis anatolicus Korge, 1964, nov.syn., = Q. troglophilus Coiffait, 1969, nov.syn.; Quedius job Coiffait, 1963 = Q. lydus Fagel, 1968, nov.syn.; Quedius humeralis Stephens, 1832 = Q. coxalis Kraatz, 1858, nov.syn., = Q. atticus Coiffait, 1967, nov.syn.; Quedius boops (Gravenhorst, 1802) = Q. haafi Scheerpeltz, 1956, nov.syn. Lectotypes are designated for Quedius cyanescens Mulsant & Rey, 1876, Q. bulgaricus Scheerpeltz, 1937, Q. albanicus Bernhauer, 1926, and Q. schipkanus Scheerpeltz, 1937. A neotype is designated for Q. josue Saulcy, 1865. The male sexual characters of several species are illustrated. Two species are redescribed. The East Mediterranean species allied to Quedius nivicola Kiesenwetter, 1858 are revised. Their zoogeography and natural history are discussed. The currently known distributions of ten species are mapped.

529:
A revision of Medon. XI. Five new species, additional recortds, and the first confirmed records from the Oriental region (Coleoptera: Staphylinidae: Paederinae). – Five species of the paederine genus Medon Stephens, 1833 from the East Palaearctic and Oriental regions are described and illustrated: Medon arunicus nov.sp. (East Nepal: Arun valley) and M. acutapicalis nov.sp. (East Nepal: Theratum district) of the M. apicalis group; M. sundaicus nov.sp. (Indonesia: Java, Sulawesi) and M. sabahicus nov.sp. (Malaysia: Sabah) of the M. ferrugineus group; M. emeianus nov.sp. (China: Emei Shan) of the M. profundus group. Medon sundaicus and M. sabahicus represent the first confirmed records of the genus from the Oriental region. Additional records of 32 species and subspecies are reported, among them several new country records. Medon is now represented in the Palaearctic region by five species groups and 86 confirmed species, 23 of which are distributed in the East Palaearctic region.

530:
On the Staphylinidae of the Greek island Corfu (Insecta: Coleoptera). – A study of nearly 10,000 specimens of Staphylinidae collected in the Ionian island Corfu, Greece, in late spring 2017 yielded more than 233 species. Additional, previously unpublished records of 66 named species are reported from the island. Two species are described and illustrated: Borboropora corcyrana Assing spec. nov. of the Aleocharinae and Ocypus corcyranus Assing spec. nov. of the Staphylininae. As many as 118 named species are reported from Corfu for the first time, 21 of these species represent first records from Greece. One name is revalidated and six names are synonymized: Euplectus jonicus Meggiolaro, 1966 (revalidated) = E. jonicus corcyreus Meggiolaro, 1966, syn. nov.; Mycetoporus punctipennis Scriba, 1868 = M. insulanus Luze, 1901, syn. nov.; Anotylus tetracarinatus (Block, 1799) = A. corcyranus (Coiffait, 1968), syn. nov.; Bledius corniger Rosenhauer, 1856 = B. bubalus Gistel, 1857, syn. nov.; Paederus littoralis Gravenhorst, 1802 = P. pelikani Reitter, 1884, syn. nov.; Leptacinus batychrus (Gyllenhal, 1827) = Phacophallus corcyranus Bordoni, 2017, syn. nov. Including reliable previous literature records and the new records reported in the present paper, 446 named species (plus additional unnamed species) are currently known from Corfu. Thus, the known fauna of this island is significantly more diverse than those of other East Mediterranean islands, including the much larger Cyprus. A checklist of the Staphylinidae fauna of Corfu is provided. Although at present 18 species and subspecies have been recorded exclusively from Corfu, most of them are unlikely to represent island endemics; three of these species are of doubtful taxonomic status.

531:
On the subterranean Staphylinidae fauna of South Spain (Coleoptera). – Staphylinidae from soil samples collected in South Spain, all of them obtained by soil washing, were examined. Seven species belonging to three subfamilies are described and illustrated: Alevonota alboculata sp. n. (Murcia) of the Aleocharinae, Lusitanopsis lencinai sp. n. (Murcia) of the Osoriinae, and Entomoculia (Stenotyphlus) solanae sp. n. (Castilla-La Mancha: Albacete), Paratyphlus deformis sp. n. (Murcia), P. magnispina sp. n. (Castilla-La Mancha: Ciudad Real), P. lencinai sp. n. (Murcia), and P. carmeloi sp. n. (Murcia) of the Leptotyphlinae. Paraleptusa anophthalma (Eppelsheim, 1884) is recorded for the first time since the original description; its habitus and the primary sexual characters are illustrated. An overview of the subterranean Staphylinidae fauna previously known from South Spain is provided. The distributions of three genera in (the south of) the Iberian Peninsula are mapped. New records of some additional species are reported.

532:
On some species of Apatetica Westwood (Coleoptera: Staphylinidae: Apateticinae). – Eight species of Apatetica Westwood, 1848 are (re-)described and illustrated, six of them for the first time: Apatetica viridipennis Fauvel, 1895 (North India); A. glaucipennis spec. nov. (Nepal); A. confusa spec. nov. (Myanmar; previously confounded with A. viridipennis); A. aspera spec. nov. (China: Sichuan); A. laevicollis Fauvel, 1904 (Vietnam); A. glabra spec. nov. (China: Yunnan); A. gibba spec. nov. (China: Gansu); A. curtipennis spec. nov. (China: Yunnan). A lectotype is designated for Apatetica laevicollis. The first confirmed record of A. laevicollis since the original description is reported. The genus currently includes 25 described species distributed in the southern East Palaearctic and Oriental regions.

533:
Five new species of Lathrobium from Nepal and China (Coleoptera: Staphylinidae: Paederinae). – Five species of the Holarctic genus Lathrobium Gravenhorst, 1802 are described and illustrated: L. adsurgens spec. nov. (Northwest Nepal: Gurans Himal) of the L. deuvei group, L. guransicum spec. nov. (Northwest Nepal: Gurans Himal) of the L. emodense group, L. latius spec. nov. (Northwest Nepal: Karnali) of the L. jumlense group, L. egens spec. nov. (China: Sichuan), a close relative of L. diffissum Assing, 2013, and L. bitaleatum spec. nov. (China: Sichuan) of the L. bibaculatum group. Lathrobium is currently represented in the Palaearctic region by as many as 612 described species, 222 of which have been recorded from China and 79 from the Himalaya.

534:
Revision of the Cousya species of the West Palaearctic Region (Coleoptera: Staphylinidae: Aleocharinae). – Species of the genus Cousya Mulsant & Rey, 1875 and five previously unrevised species of Ocyusa Kraatz, 1856 of the West Palaearctic region including Middle Asia are revised. In all, 19 species of Cousya are recognized, with one additional species of which no material was available of doubtful identity and generic affiliations. Three or four additional species remain unnamed for want of males. Only two species, O. maura (Erichson, 1837) and O. picina (Aubé, 1850), remain in Ocyusa. Four species are newly described: Cousya acris n. sp. (South Greece, South Turkey, Cyprus, Lebanon); C. struyvei n. sp. (South Spain); C. pauli n. sp. (Southwest Turkey); C. sufflata n. sp. (Greece: South Pelopónnisos). Redescriptions are provided for Cousya and ten of its species. All the Cousya species and most of the species moved to other genera are illustrated. Eleven synonymies are proposed: Cousya bicolor (Bernhauer, 1900) = C. mirabilis Assing, 2011, n. syn.; C. defecta (Mulsant & Rey, 1875) = C. humicola Fagel, 1965, n. syn.; C. longitarsis (Thomson, 1867) = C. rugipennis (J. Sahlberg, 1890) n. syn. (previously a synonym of Oxypoda funebris Kraatz, 1856), = C. peezi Scheerpeltz, 1957, n. syn.; C. nigrata (Fairmaire & Laboulbène, 1856) = C. cephallenica (Scheerpeltz, 1931), n. syn., = C. nitidiventris Fagel, 1958, n. syn., = C. lakloukensis Fagel, 1965, n. syn.; C. schuelkei Assing, 2007 = C. planicollis Assing, 2011, n. syn.; C. bimaculata (Fauvel, 1899) = C. vaulogeri (Bernhauer, 1936), n. syn.; Oxypoda flavicornis Kraatz, 1856 = Ocyusa beieri Scheerpeltz, 1931, n. syn.; Ocyusa pellax Peyerimhoff, 1919 = Oxypoda argus Normand, 1935, n. syn. Six new binomina are established: Cousya dissoluta (Eppelsheim, 1888), n. comb. (ex Ocyusa); C. praecox (Eppelsheim, 1888), n. comb. (ex Ocyusa); Oxypoda picta Mulsant & Rey, 1875, n. comb. (ex Cousya); Oxypoda pellax (Peyerimhoff, 1919), n. comb. (ex Ocyusa); Oxypoda heydeni (Eppelsheim, 1879), n. comb. (ex Ocyusa); Tectusa uhligi (Pace, 1987), n. comb. (ex Cousya). The material labelled as types of Cousya uhligi is composed of three species, all of them belonging to Tectusa Bernhauer, 1899. One of them is Tectusa pirinica n. sp. (Bulgaria: Pirin Planina). Four Bulgarian species of Tectusa, including T. pirinica, are illustrated. “Ocyusa” apicalis Normand, 1935 is not congeneric with the type species of Ocyusa and treated as Oxypodina incertae sedis. A neotype is designated for Ocyusa defecta Mulsant & Rey, 1875. Lectotypes are designated for Cousya peezi Scheerpeltz, 1957, Ocyusa eppelsheimi Bernhauer, 1902, O. araxis Bernhauer, 1902, O. bicolor Bernhauer, 1900, Homalota dissoluta Eppelsheim, 1888, H. praecox Eppelsheim, 1888, and Ocyusa ferdinandicoburgi Rambousek, 1909. A catalogue and a key to the Cousya species of the West Palaearctic region are provided. The species of Cousya are assigned to two species groups, the C. nigrata group (eleven species) and the C. crocea group (eight species). While the species of the C. nigrata group are fully winged and widespread, those of the C. crocea group are partly incapable of flight and have restricted distributions; six of them have been recorded only from their respective type localities. The distributions of all the Cousya species are mapped. Available evidence suggests that at least the species of the C. nigrata group reproduce in autumn (most species in a subterranean habitat), have their preimaginal development in winter and early spring, emerge from the pupa and disperse in spring, and spend the warmer seasons either in estivation or in a subterranean habitat. A biased sex ratio (males rarer than females) was observed in two species. It is particularly pronounced in C. nigrata: only 15 % of the sexed specimens are males. Not a single male was found in the north of its range, suggesting that the species is parthenogenetic in this region.

535:
An enigmatic new genus and species of Anthophagini from Georgia (Coleoptera: Staphylinidae: Omaliinae). – Caucanthobium nov.gen. mirandum nov.sp., an enigmatic new genus and species from Southwest Georgia, is described, illustrated, and distinguished from Anthobium Leach, 1819 and Deinopteroloma Jansson, 1947. Its currently known distribution is mapped.

536:
A revision of Palaearctic and Oriental Scymbalium and Micrillus III. New species, new combinations, and additional records (Coleoptera: Staphylinidae: Paederinae). – Six species of Scymbalium Erichson, 1839 and Micrillus Raffray, 1873 are described and illustrated: Scymbalium rossii nov.sp. (Cambodia), Micrillus virgatus nov.sp. (Cambodia, South Thailand) of the newly established M. badius group, M. hamatus nov.sp. (South Morocco) of the M. testaceus group, M. kongi nov.sp. (Cambodia) of the M. suturalis group, and M. coloratus nov.sp. (Cambodia) and M. rougemonti nov.sp. (Sri Lanka) of the newly established M. coloratus group. Based on a re-assessment of the inter- and intrageneric phylogenetic affiliations of Scymbalium and Micrillus, two species are moved from Scymbalium to Micrillus: Micrillus badius (Motschulsky, 1858), nov.comb.; Micrillus nepalensis (Assing, 2013), nov.comb. Additional records of one species of Scymbalium and of ten species (one of them undescribed) of Micrillus are reported, among them five first records from Cambodia (3), Syria (1), and Indonesia (1).

537:
A revision of Pinobius. II. Three new species from Cambodia and Indonesia, and additional records (Coleoptera: Staphylinidae: Paederinae). – Three species of the dolicaonine genus Pinobius Macleay, 1871 are described and illustrated: Pinobius acutus nov.sp. (Cambodia) and P. rossii nov.sp. (Cambodia) of the P. indicus group, and P. forcipifer nov.sp. (Indonesia: Kalimantan Timur) of the P. major group. Additional records of six previously described species are reported, among them a new record from Cambodia. The genus now includes a total of 38 described species, 30 of which are represented in the southern Palaearctic, Oriental, and Australian regions.

538:
A revision of Palaearctic and Oriental Pseudolathra. V. Two new species from Cambodia and Thailand, and additional records (Coleoptera: Staphylinidae: Paederinae). – SPseudolathra armigera nov.sp. (Cambodia) and P. aviformis nov.sp. (North Thailand) are described and illustrated. One synonymy is proposed: Pseudolathra fissa Assing, 2013 = P. ceylonica Rougemont, 2015, nov.syn. Additional records of seven previously described species are reported, among them several new country records. The genus is reported from Cambodia (four species) and Bhutan for the first time and currently represented in the East Palaearctic and Oriental regions by 40 described species. A catalogue of these species is provided.

539:
On some Myllaena species in the East Mediterranean and Caucasus regions (Coleoptera: Staphylinidae: Aleocharinae). – Two species of previously uncertain status of Myllaena Erichson, 1837 are revised, redescribed, and illustrated: M. graeca Kraatz, 1858 (Greece, Turkey) and M. caucasica Eppelsheim, 1880 (Caucasus region). A neotype and a lectotype are designated for M. graeca and M. caucasica, respectively. Three species are described and illustrated: M. mutabilis nov.sp. (Greece: Corfu), M. cretica nov.sp. (Greece: Crete), and M. ambulans nov.sp. (Armenia). Eight first records of Myllaena species from Armenia (3), Nagorno-Karabakh (2), Russia (1), Azerbaijan (1), and Iran (1) are reported. The currently known distributions of M. graeca, M. caucasica, and of the three newly described species are mapped.

540:
A revision of Geostiba of the West Palaearctic region. XXVI. New species and additional records, primarily from the Caucasus region (Coleoptera: Staphylinidae: Aleocharinae). – Seven species of Geostiba Thomson, 1858 from the Caucasus region and Ukraine are described and illustrated: Geostiba (Tropogastrosipalia) reducta nov.sp. (Russia: West Caucasus); G. (T.) defecta nov.sp. (Russia: West Caucasus); G. (Sibiota) breviflagellata nov.sp. (Georgia: Imereti); G. (S.) nigrohortensia nov.sp. (Nagorno-Karabakh); G. (S.) deliqua nov.sp. (South Armenia); G. (Sipalotricha) marmotae nov.sp. (Ukraine); G. (S.) manca nov.sp. (Georgia: Imereti). The primary sexual characters of several previously described species are illustrated. Geostiba cingulata (Eppelsheim, 1878), previously in the subgenus Chondridiosipalia Scheerpeltz, 1951 is moved to the subgenus Sipalotricha Scheerpeltz, 1951. Additional records of eight Caucasian species are reported. The currently known distributions of 15 Caucasian species of the subgenera Tropogastrosipalia Scheerpeltz, 1951, Sibiota Casey, 1906, and Sipalotricha are mapped..

541:
A new genus of Oxypodini from China (Coleoptera: Staphylinidae: Aleocharinae). – Notocousya nov.gen. quadriceps nov.sp. (China: Yunnan) of the Oxypodini, subtribe Oxypodina, is described and illustrated. The new genus currently includes only the type species N. quadriceps and is characterized by a distinctive morphology of the head, including the mouthparts, and of the pronotum. The absence of previous records suggests a cryptic reproduction habitat.

542:
Revision of the Baeoglena species of the West Palaearctic Region (Coleoptera: Staphylinidae: Aleocharinae). – The subgenus Baeoglena Thomson, 1867 of the speciose aleocharine genus Oxypoda Mannerheim, 1830 has been subject to considerable taxonomic confusion rendering a reliable identification of material from regions other than Central Europe and the Canary Islands virtually impossible. Based on a revision of abundant material from various major public and private collections, seven species are distributed in the West Palaearctic region exclusive of the Canary Islands. Two new species are described and illustrated: Oxypoda (Baeoglena) rectacia spec. nov (East Mediterranean, from South Greece to the Middle East) and O. (B.) derecta spec. nov. (West Caucasus, Northeast Anatolia). Diagnoses and illustrations of the genitalia are provided for the remaining five species. The following synonymies are established: Oxypoda nova Bernhauer, 1902 = O. giachinoi Pace, 2001, syn. nov.; O. hispanica Fagel, 1958 and its replacement name O. inexpectata Fagel, 1965 are removed from synonymy with O. praecox Erichson, 1839 and synonymized with O. fusina Mulsant & Rey, 1875. Oxypoda recondita Kraatz, 1856 and O. dalmatina Bernhauer, 1905 are excluded from Baeoglena and moved to the subgenus Bessopora Thomson, 1859. Lectotypes are designated for Oxypoda nova Bernhauer, 1902, O. caucasica Bernhauer, 1902, and O. kuehnelti Scheerpeltz, 1963. An outline of the taxonomic history, a diagnosis of the subgenus, a checklist, and a key to species are provided. A zoogeographic analysis revealed some remarkably discontinuous distributions and parapatric distribution patterns which are plausibly explained only with interspecific competition among Baeoglena species. The distributions of the subgenus as a whole and of the individual species in the West Palaearctic region are mapped.

543:
On the Lomechusini fauna of the Palaearctic and Oriental regions. XXVI. New species, a new synonymy, and additional records (Coleoptera: Staphylinidae: Aleocharinae). – Types and additional material of twelve genera of Lomechusini from the Palaearctic and Oriental regions are revised. In all, 24 species are described and/or illustrated, 21 of them new: Pella feldmanni spec. nov. (China: Sichuan) of the P. cognata group; Orphnebius fortesetosus spec. nov. (China: Sichuan) and O. taurus spec. nov. (Cambodia) of the O. hauseri group; Orphnebius (Deroleptus) morulus spec. nov. (South Vietnam), O. (D.) globulifer spec. nov. (Malaysia: Pahang), O. (D.) elevatus spec. nov. (Laos), and O. (D.) moruliflavus spec. nov. (Malaysia: Sabah) of the O. siwalikensis group; Keratodegnathus cornutus spec. nov. (Malaysia: Pahang); Amaurodera schmidti spec. nov. (East Nepal) of the A. silvana group; A. brevipennis spec. nov. (Malaysia: Sabah); A. atra spec. nov. (Philippines: Mindanao); Drusilla bispinosa spec. nov. (Malaysia: Pahang); D. shavrini spec. nov. (Philippines: Mindanao); D. penicillata spec. nov. (Philippines: Mindanao); D. spiniventris spec. nov. (Philippines: Mindanao); D. breviuter spec. nov. (Philippines: Mindanao); Pheidologitonetes biplicatus spec. nov. (China: Yunnan); Zyras (Zyras) conlectus spec. nov. (Myanmar); Z. (Z.) hetzeli spec. nov. (Taiwan); Z. (Z.) reductus spec. nov. (Philippines: Mindanao); Z. (Z.) shavrini spec. nov. (Philippines: Mindanao). The following synonymy is established: Drusilla obliqua (Bernhauer, 1916) = D. zyrasoides Dvořák, 1988, syn. nov. Additional records of 49 named species are reported, among them twelve first records from Greece (1), Lebanon (1), Israel (1), Jordan (1), Iran (1), Nepal (1), China (1), Taiwan (1), Laos (1), Cambodia (2), and Borneo (1). The distribution of Myrmoecia urartu (Iablokoff-Khnzorian, 1962) is mapped.

544:
The Staphylinidae of Armenia and Nagorno-Karabakh (Coleoptera). – The previously largely neglected and poorly known staphylinid faunas of Armenia and Nagorno-Karabakh are addressed. Based on a study of more than 31,000 Staphylinidae recently collected in various habitats and using different methods, and on a critical evaluation of previous literature records, a checklist of the faunas of Armenia and Nagorno-Karabakh is compiled. The fauna of Armenia currently includes 675, that of Nagorno-Karabakh 198 named species. Nevertheless, it is concluded that the species inventory of both regions, especially that of Nagorno-Karabakh, is still far from complete. As many as 262 and 183 species are reported from Armenia and Nagorno-Karabakh, respectively, for the first time. A list of 99 species erroneously or doubtfully recorded from the study region is provided. A comparison with the species number and systematic composition of the faunas of other Caucasian countries and regions revealed that (a) their known diversities are significantly lower than should be expected and (b) a remarkably high proportion (nearly 40 %) of Aleocharinae in the faunas of Armenia and Nagorno-Karabakh, with the genus Atheta Thomson, 1858 alone accounting for approximately 10 % of the total diversity in Armenia. The faunas of Armenia and Nagorno-Karabakh are primarily composed of widespread species, many of them Caspian (Caucasian) and Iranian elements, and remarkably few endemics. Only 23 species, thirteen of the Aleocharinae (most of them belonging to the genus Geostiba Thomson, 1858), four of the Pselaphinae, five of Scydmaeninae, and one of Staphylininae are classified as regional endemics, and one species of Aleocharinae from the peak region of Mount Khustup is classified as a local endemic. The distribution of one addititional species is confined to South Armenia and adjacent parts of North Iran. A number of species is currently known only from Armenia, but of doubtful taxonomic status or unlikely to represent endemics. Records of some species in Armenia and/or Nagorno-Karabakh revealed some remarkably discontinuous distributions with gaps of up to approximately 2,800 km; three of these distributions are mapped. Fourteen species are newly described: Omalium kociani Zanetti spec. nov. (Armenia: Jermuk) of the Omaliinae, Proteinus baculatus Assing spec. nov. (Armenia; Northeast Turkey) of the Proteininae, Bryaxis armeniacus Brachat spec. nov. (Armenia: NW Hrazdan) and B. meghruicus Brachat spec nov. (South Armenia: Meghru range) of the Pselaphinae, Atheta (Paralpinia) meghruica Assing spec. nov. (South Armenia: Meghru range), Bellatheta khustupica Assing spec. nov. (South Armenia: Mount Khustup), Calodera alticola Assing spec. nov. (Armenia: Mount Karkar), and Tachyusa unguis Assing spec. nov. (South Armenia) of the Aleocharinae, Anotylus hamatoides Schülke spec. nov. (Armenia) of the Oxytelinae, Euconnus (Tetramelus) longilaminatus Meybohm spec. nov. (North Armenia), E. (T.) tavushus Meybohm spec. nov. (North Armenia), E. (T.) karabakhus Meybohm spec. nov. (Nagorno-Karabakh), Neuraphes (Paraphes) gomarantsus Meybohm spec. nov. (South Armenia: Meghru range), and N. (P.) syunikus Meybohm spec. nov. (South Armenia) of the Scydmaeninae. Eight synonymies and one revalidation are established: Dialycera minuta Luze, 1906 = Phyllodrepa armena Iablokoff-Khnzorian, 1959, syn. nov.; Mycetoporus silvaticus Iablokoff-Khnzorian, 1962 = M. dispersus Schülke & Kocian, 2000, syn. nov.; Aleochara subtumida (Hochhuth, 1859) = Aleochara khnzoriani Amiryan, 1999, syn. nov.; Platystethus cephalotes Eppelsheim, 1878, revalidated (previously synonym of P. laevis Märkel & Kiesenwetter, 1848) = P. oblongopunctatus Roubal, 1911, syn. nov.; Euconnus lalvarensis Iablokoff-Khnzorian, 1964 = Euconnus pseudorobustus Franz, 1986, syn. nov.; Astenus rufopacus Reitter, 1909 = A. baali Coiffait, 1960, syn. nov.; Heterothops dissimilis (Gravenhorst, 1802) = H. armeniacus Coiffait, 1977, syn. nov.; Heterothops praevius Erichson, 1839 = Heterothops montanus Iablokoff-Khnzorian, 1966, syn. nov.

545:
A revision of Palaearctic and Oriental Rugilus. VI. Two new species from China, a new synonymy, and additional records (Coleoptera: Staphylinidae: Paederinae). – Two species of Rugilus Leach, 1819 from China are described and illustrated: R. (Rugilus) reuteri nov.sp. (North Sichuan) and R. (R.) gryps nov.sp. (West Sichuan: Gongga Shan). Additional records of 27 previously described species, 18 of the nominal subgenus, seven of Eurystilicus Fagel, 1853, and two without subgeneric assignment, are reported from the Palaearctic and Oriental regions, among them six new country records from Pakistan (3), Lebanon (1), India (1), and Cambodia (1). A new synonymy is proposed: Rugilus arabs (Saulcy, 1865) = R. couloni (Drugmand, 1989), nov.syn. The preoccupied name Rugilus rugossimus Assing, 2015 is replaced with R. asperrimus nov.nom. Stilicus ovicollis MacLeay, 1873, a species erroneously transferred to Rugilus recently, is reassigned to Scopaeus Erichson, 1839. New illustrations of the male sexual characters of R. prodoni (Coiffait, 1982), R. bagmaticola Rougemont, 1998, and R. curvatus Assing, 2013 are provided. The distributions of the species of the R. malaisei group are mapped. Including the new species, Rugilus is now represented in the Palaearctic and Oriental regions by 102 species and one subspecies; 35 species are currently known from China.

546:
A revision of Palaearctic and Oriental Scymbalium and Micrillus IV. The fauna of Cambodia (Coleoptera: Staphylinidae: Paederinae). – Records of one species of Scymbalium Erichson, 1839 and seven species of Micrillus Raffray, 1873 are reported from Cambodia. Three of them are described and illustrated: Micrillus rossianus nov.sp. of the M. badius group, M. variceps nov.sp. of the newly established M. variceps group, and M. bispinosus nov.sp. of the M. coloratus group. Including these additions, the Cambodian fauna is currently composed of one Scymbalium and nine Micrillus species and thus more diverse than that of any other country in the Oriental region.

547:
Three new species and additional records of Oedichirus (Coleoptera: Staphylinidae: Paederinae). – Three species of Oedichirus Erichson, 1839 are described and illustrated: O. pauli nov.sp. (North Vietnam), O. puicus nov.sp. (North Thailand), and O. russipennis nov.sp. (Indonesia: Papua: Yapen). A lectotype is designated for O. terminatus Erichson, 1843, a species described from Angola and subsequently reported also from the Mediterranean region. The sexual characters of O. terminatus are illustrated. Additional records of three species are provided.

548:
On the taxonomy of some West Palaearctic Quedius species, with descriptions of new species and new synonymies (Coleoptera: Staphylinidae: Staphylininae). – Three species of Quedius Stephens, 1829 are described and illustrated, two of them new: Quedius (Raphirus) iridicolor Quedenfeldt, 1882, Q. (Microsaurus) morulus nov.sp. (Iran), and Q. (M.) limans nov.sp. (Israel). Three synonymies are established: Quedius umbrinus Erichson, 1839 = Q. kuboni Štourač, 1998, nov.syn.; Q. cohaesus Eppelsheim, 1888 = Q. pseudonigriceps Reitter, 1909, nov.syn.; Q. cruentus (Olivier, 1795) = Q. xanthurus Iablokoff-Khnzorian, 1961, nov.syn. Diagnoses distinguishing Q. iridicolor and Q. nigriceps Kraatz, 1857, as well as Q. (Microsaurus) invreae Gridelli, 1925 and Q. (M.) puncticollis (Thomson, 1867) are provided. Previous erroneous records of Q. puncticollis are rectified. Quedius iridicolor is re-assigned to the subgenus Raphirus Stephens, 1829. The currently known distribution of Q. iridicolor is mapped.

549:
A revision of the Alevonota species of the Palaearctic region. III. Two new species from Taiwan, a new combination, a new synonymy, and additional records (Coleoptera: Staphylinidae: Aleocharinae). – Alevonota calliceroides nov.sp. and A. hetzeli nov.sp., both of them from the same locality in Hualien Hsien, Taiwan, are described and illustrated. Homalota hepatica Erichson, 1839, the type species of Enalodroma Thomson, 1859, is transferred to Alevonota Thomson, 1858, rendering Enalodroma a junior synonym of Alevonota and yielding the binomen Alevonota hepatica (Erichson, 1839), nov.comb. Data on the natural history of A. hepatica are compiled. Additional records of four species are reported, among them several new country records of A. hepatica and the first record of A. gracilenta (Erichson, 1839) from Ukraine.

550:
Five new species of Chinecallicerus from China (Coleoptera: Staphylinidae: Aleocharinae: Geostibini). – Five species of Chinecallicerus Assing, 2004 from China are described and illustrated: Chinecallicerus formidabilis nov.sp. (North Sichuan), C. glabriventris nov.sp. (West Yunnan), C. reuteri nov.sp. (North Sichuan), C. feldmanni nov.sp. (North Sichuan), and C. discrepans nov.sp. (North Sichuan). An updated key to the Chinecallicerus species is provided. The distributions of the new species and their respective closest relatives are mapped. Including the new species, the genus now includes 18 species, nine of which have been recorded from the Chinese province Yunnan, eight from Sichuan, and one from Qinghai. All the species are known only from their respective type localities, fourteen of them are represented exclusively by their respective holotypes.

551:
A revision of the species of “Blepharhymenus” of the Palaearctic and Oriental regions (Coleoptera: Staphylinidae: Aleocharinae: Oxypodini). – Species of Blepharhymenus Solier, 1849 had been reported from the Chilean Subregion (including the type species of the genus), the Nearctic, Palaearctic, Ethiopian, and Australian regions, suggesting that the overall distribution of this genus was remarkably discontinuous. In order to clarify the status and generic assignment of the Palaearctic species, types and additional material are revised, including also a selection of species from Chile and North America. A study of this material revealed that what had been regarded as Blepharhymenus in fact represents various distinct lineages and that none of the species from the Palaearctic Region belongs to the clade containing the type species of Blephar­hymenus. The following genus group taxa are newly described, revalidated, or elevated to genus: Echidnoglossa Wollaston, 1864, revalidated (previously synonym of Blepharhymenus); Maur­echidna subgen.n. (subgenus of Echidnoglossa; type species: Echidnoglossa ventricosa Queden­feldt, 1881); Orechidna subgen.n. (subgenus of Echidnoglossa; type species: Echidnoglossa hirthei sp.n.); Sinechidna subgen.n. (subgenus of Echidnoglossa; type species: Blepharhymenus smetanai Pace, 2012); Syntomenus Bernhauer, 1939, stat.n. (previously subgenus of Blepharhymenus); Kortomenus gen.n. (type species: Blepharhymenus koreanus Paśnik, 2001); Colusa Casey, 1885, re­validated (previously synonym of Blepharhymenus; type species Colusa gracilis Casey, 1885). Echidnoglossa is discontinuously distributed across the Palaearctic and northern Oriental regions and currently includes 19 species in five subgenera. Syntomenus and the monotypical genus Kortomenus are confined to the southern East Palaearctic and northern Oriental regions. All the species from the Palaearctic and Oriental regions are (re-)described and illustrated, six of them for the first time: Echidnoglossa (Echidnoglossa) russa sp.n. (Egypt: Sinai Peninsula); Echidnoglossa (Orechidna) hirthei sp.n. (Nepal); E. (O.) betzi sp.n. (Thailand); E. (O.) artior sp.n. (Taiwan); E. (O.) formosana sp.n. (Taiwan); Syntomenus laoticus sp.n. (Laos). Numerous new combinations are proposed: the Pa­laearctic species previously attributed to Blepharhymenus are all assigned to Echidnoglossa, Syn­tomenus, or Kortomenus, and the binomen Ocalea dabensis (Pace, 2012), comb.n. (ex Blephar­hymenus) is established. The Nearctic species are tranferred to Colusa. The following synonymies are proposed: Echidnoglossa corsica Mulsant & Rey, 1875 = E. sardoa (Scheerpeltz, 1954), syn.n.; E. glabrata (Kiesenwetter, 1870) = E. moczarskii (Scheerpeltz, 1954), syn.n.; E. maghrebica (Fagel, 1960) = E. peyerimhoffi (Fagel, 1960), syn.n.; E. meschniggi (Bernhauer, 1936) = E. ele­gans (Fagel, 1959), syn.n. = E. guadalupensis (Fagel, 1959), syn.n.; Syntomenus chinensis (Bern­hauer, 1939) = S. rougemonti (Pace, 1999), syn.n. Lectotypes are designated for Blephar­hy­menus moczarskii Scheerpeltz, 1954, Echidnoglossa paulinoi Skalitzky, 1884, and E. ventri­cosa Que­denfeldt, 1881. A checklist of the taxa distributed in the Palaearctic and Oriental regions and a key to the species of Echidnoglossa is provided. The distributions of Echidnoglossa species are mapped.

552:
Three new species of Platyprosopus from Thailand, Cambodia, and Sierra Leone, and additional records (Coleoptera, Staphylinidae, Staphylininae). – Three species of Platyprosopus Mannerheim, 1830 are described and illustrated: P. latitarsalis nov.sp. (Thailand); P. rossii nov.sp. (Cambodia); P. maximus nov.sp. (Sierra Leone). Additional records of three species are reported primarily from Cambodia, among them two new country records.

553:
A revision of Geostiba of the West Palaearctic region. XXVII. New species from Georgia and Kyrgyzstan, and additional records (Coleoptera, Staphylinidae, Aleocharinae). – Five species of Geostiba Thomson, 1858 from Georgia and East Kyrgyzstan are described and illustrated: Geostiba (Tropogastrosipalia) simplicicollis nov.sp. (Georgia: Kakheti); G. (Sibiota) kakhetiana nov.sp. (Georgia: Kakheti); G. (S.) frischi nov.sp. (Kyrgyzstan: Issyk-Kul); G. (S.) prominens nov.sp. (Kyrgyzstan: Issyk-Kul); G. (S.) dentata nov.sp. (Kyrgyzstan: Issyk-Kul). Based on a study of recently collected material, Geostiba zerchei Pace, 1996 is moved from the subgenus Sibiota Casey, 1906 to Sipalotricha Scheerpeltz, 1931; the male sexual characters of this previously doubtful taxon are described and illustrated for the first time. Additional records of eight species are reported from the Caucasus region.

554:
The mother of synonyms: on the Meotica species of the Palaearctic Region (Coleoptera, Staphylinidae, Aleocharinae, Oxypodini). – The Holarctic genus Meotica Mulsant & Rey, 1873 has been in a state of serious taxonomic confusion, primarily because numerous species have been described without a study of the sexual characters and because both intraspecific variation and distribution ranges have been underestimated. Based on a revision of types and additional material, the genus is represented in the Palaearctic region by 20 species, 18 of them distributed in the West Palaearctic and two in the East Palaearctic regions. The identity of three additional species described from Northwest Africa, whose type material is currently inaccessible, remains uncertain. One genus-group and as many as 23 species-group synonymies are proposed for the first time: Meotica Mulsant & Rey, 1873 = Cryptusa Mulsant & Rey, 1873, nov.syn.; Meotica filiformis (Motschulsky, 1860) = M. capitalis Mulsant & Rey, 1873, nov.syn., = M. fageli Benick, 1953, nov.syn.; M. exilis (Gravenhorst, 1806) = M. simillima Benick, 1953, nov.syn.; M. pallens (Redtenbacher, 1849) = M. hanseni Benick, 1953, nov.syn.; M. filaria (Fauvel, 1898) = M. meridiogallica Scheerpeltz, 1954, nov.syn.; M. parasita Mulsant & Rey, 1873 = M. winkleri Benick, 1953, nov.syn., = M. bosnica Scheerpeltz, 1954, nov.syn., = M. albanicola Scheerpeltz, 1969, nov.syn., = M. aegyptia Pace, 1983, nov.syn.; M. moczarskii Scheerpeltz, 1927 = M. finnmarchica Benick, 1953, nov.syn., = M. romana Benick, 1953, nov.syn., = M. albanica Benick, 1953, nov.syn., = M. marchica Benick, 1953, nov.syn., = M. exigua Benick, 1953, nov.syn., = M. alpina Benick, 1953, nov.syn., = M. testacea Benick, 1953, nov.syn., = M. neapolitana Scheerpeltz, 1954, nov.syn., = M. stockmanni Muona, 1978, nov.syn., = M. anglica Benick, 1991, nov.syn.; M. ochsi Benick, 1953 = M. punctulata Benick, 1953, nov.syn.; M. normandi Benick, 1953 = M. franzi Pace, 1983, nov.syn.; M. caucasica Benick, 1953 = M. decolor Assing, 2004, nov.syn.; Apimela sabulicola (Bernhauer, 1914) = Meotica quadraticollis Scheerpeltz, 1954, nov.syn. Thus, Meotica now totals 58 synonyms outnumbering the valid names by a factor of nearly three. The five most widespread species alone account for 53 synonyms: M. moczarskii (19 synonyms), M. filiformis (12), M. exilis (8), M. pallens (8), and M. parasita (6). Meotica kuehnelti (Scheerpeltz, 1963) is transferred to the genus Tectusa Bernhauer, 1899. Since Tectusa kuehnelti (Scheerpeltz, 1963) is preoccupied by T. kuehnelti (Scheerpeltz, 1962), the name is replaced with Tectusa killinica nov.nom. Lectotypes are designated for Meotica hoelzeli Benick, 1953, M. clavata Benick, 1953, M. simillima Benick, 1953, M. bosnica Scheerpetz, 1954, M. albanica Scheerpetz, 1967, M. moczarskii Scheerpeltz, 1927, M. marchica Benick, 1953, M. alpina Benick, 1953, M. exigua Benick, 1953, M. roubali Benick, 1953, M. neapolitana Scheerpeltz, 1954, M. ochsi Benick, 1953, M. smetanai Scheerpetz, 1967, M. normandi Benick, 1953, M. quadraticollis Scheerpeltz, 1954, and Atheta filaria Fauvel, 1898. A synonymic catalogue, as well as concise diagnoses and illustrations of the sexual characters of all the species of which material was available are provided. 

555:
New species and additional records of Ischnosoma from Georgia (Coleoptera, Staphylinidae, Tachyporinae). – Four species of Ischnosoma Stephens, 1829 from Georgia are described and illustrated, all of them belonging to the I. spelaeum group: Ischnosoma acre nov.sp. (Southwest Georgia: Imereti region), a close relative of I. solodovnikovi Schülke, 2001 and I. molle Assing & Schülke, 2017, I. convergens nov.sp. (Southwest Georgia: Adjara region: Meskheti Range) and I. parallelum nov.sp. (Southwest Georgia: Adjara region: Shavsheti Range), both closely allied to I. myops (Eppelsheim, 1880), and I. cuspidatum nov.sp. (Northeast Georgia: Kakheti region). For comparison, new illustrations of I. solodovnikovi, I. molle, and I. myops are provided. Additional records of four species are reported, among them the first confirmed record of I thoracicum (Eppelsheim, 1880), a species with a most likely erroneous type locality. The distributions of the new species and of I. solodovnikovi, I. molle, I. myops, and I. thoracicum are mapped.

556:
Four new species and additional records of Lomechusini from Mindanao, Philippines (Coleoptera, Staphylinidae, Aleocharinae). – Drusilla bucina nov.sp., D. holoserica nov.sp., Zyras (Zyras) rasilis nov.sp., and Z. (Z.) nigrorufus nov.sp. (all Philippines: Mindanao) are described and illustrated. Six additional species, four of them unnamed, are reported from Mindanao. Including the new species, 46 named species of Lomechusini are currently known from the Philippines.

557:
On the Staphylinidae of the Greek island Samothraki (Insecta, Coleoptera). – Aside from individual records of two widespread species, the Staphylinidae of Samothraki, a Greek island in the north of the Aegean Sea, had never been studied. In April 2019, a field trip was conducted to Samothraki to investigate the staphylinid fauna. A study of 1372 specimens collected during this field trip yielded 123 species, 114 of them named and six of them endemic. Two species and one subspecies are reported from Greece for the first time. Seven species are described and illustrated: Apimela procera Assing nov.sp. (Greece: Samothraki, Fthiotis; Turkey: Osmaniye) of the Aleocharinae; Cyrtotyphlus samothracicus Assing nov.sp. (endemic) and Metrotyphlus samothracicus Assing nov.sp. (endemic) of the Leptotyphlinae; Cephennium samothracicum Assing nov.sp. (endemic) and Scydmoraphes samothracicus Meybohm nov.sp. (endemic) of the Scydmaeninae; Bryaxis samothracicus Brachat nov.sp. (endemic) and Tychobythinus assingi Brachat nov.sp. (endemic) of the Pselaphinae. A checklist of the species recorded from Samothraki is provided. Samothraki is characterized by a relatively poor forest fauna, but a high diversity of wetland species. It is compared to that of the North Aegean Islands Chios, Lesbos, Samos, and Ikaría regarding total diversity and number of endemic species. In an appendix, Bryaxis pangeonicus Brachat nov.sp. from Oros Pangéo (Northeast Greece) is described.

558:
A new genus of Oxypodina from Kyrgyzstan (Coleoptera, Staphylinidae, Aleocharinae, Oxypodini). – Typhlotectusa nov.gen. frischi nov.sp. (East Kyrgyzstan: Issyk-Kul) of the subtribe Oxypodina is described and illustrated. The new species was discovered in a high-altitude habitat and is characterized by a slender habitus, its mouthparts, strongly incrassate antennae, a short metatarsomere I, the shapes of the male sternite VIII and the apical lobe of the paramere, as well as additionally by the complete absence of eyes, reduced wings, and reduced pigmentation.

559:
The first record of the subfamily Osoriinae from the Caucasus region (Coleoptera: Staphylinidae). – Leptotyphlopsis shavshetica sp. nov. from the Shavsheti Range (Southwest Georgia: Adjara region), the first representative of the subfamily Osoriinae and the first truly endogean species of Staphylinidae to be recorded from the Caucasus region, is described and illustrated. The distribution of this species is mapped.

560:
On some Proteinus species of the Caucasus region, with description of a new species from Georgia and a new synonymy (Coleoptera: Staphylinidae: Proteininae). – Proteinus cavatus sp. nov. (Southwest Georgia) of the P. longicornis group, a species with conspicuous male secondary sexual characters, is described and illustrated. Based on a study of type material, redescriptions and illustrations are provided for Proteinus planicollis Reitter, 1905 and P. reflexicollis Reitter, 1905 and lectotypes are designated for both species. A new synonymy is proposed: P. atomarius Erichson, 1840 = P. hiburnus Gistel, 1857, syn. nov. The presence of two widespread species in Georgia is confirmed.

561:
A new species of Parabolitobius from the Meskheti Range, Southwest Georgia, with a new record of Bolitobius insignis (Coleoptera: Staphylinidae: Tachyporinae). – Parabolitobius meskheticus sp. nov. (Southwest Georgia: Meskheti Range), a micropterous and presumably endemic species, is described, illustrated, and distinguished from allied and geographically close spccies. Including the new species, Parabolitobius Li, Zhao & Sakai, 2000 is currently represented in the Palaearctic region by eleven species. A new record of the rare Bolitobius insignis is reported from Southwest Georgia.

562:
On the taxonomy and zoogeography of some Xantholinus species of the Caucasus region, with descriptions of a new species from Georgia and two new synonymies (Coleoptera: Staphylinidae: Staphylininae). – Xantholinus (Calolinus) mesheticus sp. nov. (Southwest Georgia: Adjara: Mesheti Range), a micropterous species with a presumably restricted distribution, is described and illustrated. Two synonymies are proposed: Xantholinus (Helicophallus) variabilis Hochhuth, 1851 = X. kirschenblati Bordoni, 1975, syn. nov., = X. ceviki Anlaş, 2014, syn. nov. New records of five species of Xantholinus Dejean, 1821 are reported from Georgia. The distributions of six Caucasian species are mapped.

563:
On the Leptusa fauna of the Caucasus region II. A new species and additional records from Georgia, and three new synonymies (Coleoptera: Staphylinidae: Aleocharinae). – Leptusa (Stictopisalia) shavshetica sp. nov. (Southwest Georgia: Adjara: Shavsheti Range), is described, illustrated, and distinguished from similar species. Three synonymies are proposed: Leptusa (Roubaliusa) trapezuntis Pace, 1983 = L. giresunensis Sert, Turan & Kabalak, 2019, syn. nov.; Leptusa (Neopisalia) microphthalma Reitter, 1887 = L. circassica Bernhauer, 1935, syn. nov.; Leptusa (Stictopisalia) subcaucasica Pace, 1983 = L. pseudocaucasica Pace, 1983, syn. nov. Additional records of six species of Leptusa Kraatz, 1858 are reported from Georgia. The distributions of three species are mapped. The Caucasian Leptusa fauna currently includes 38 valid species in five subgenera.

564:
A revision of Othiini. XXI. Seven new species, a new synonymy, and additioonal records, with a review of the Othius fauna of Georgia (Coleoptera: Staphylinidae: Staphylininae). – Othius derectus sp. nov. (Southwest Georgia: Meskheti Range), O. unculatus sp. nov. (West Georgia: Kutaisi env.), O. rasus sp. nov. (Southwest Georgia: Meskheti Range), O. hetzeli sp. nov. (Taiwan), O. brunkei sp. nov. (North Vietnam), O. procerus sp. nov. (North Vietnam), O. devians sp. nov. (North Vietnam), and O. bispinosus sp. nov. (China: Zhejiang) are described, illustrated, and distinguished from allied species. One synonymy is proposed: Othius lapidicola Märkel & Kiesenwetter, 1848 = O. piceus Scriba, 1870, syn. nov. Additional records of twelve species are reported. The Othius fauna of Georgia currently includes 16 described species. The distributions of the Caucasian species recorded from Georgia and of the widespread, but discontinuously distributed Othius lapidicola are mapped. Othius Stephens, 1829 now comprises a total of 137 species and subspecies.

565:
New species and records of Lathrobium, with the first and southernmost record of the genus from Vietnam (Coleoptera: Staphylinidae: Paederinae). – Six species of the Holarctic genus Lathrobium Gravenhorst, 1802 are described and illustrated: L. austere sp. nov. (North Vietnam: Cao Băng Prov.); L. tsukubanum sp. nov. (Japan: Honshu: Ibaraki Pref.); L. converrens sp. nov. (Japan: Shikoku: Ehime Pref.); L. aperiens sp. nov. (Japan: Shikoku: Ehime Pref.); L. toonicum sp. nov. (Japan: Shikoku: Ehime Prov.); L. bigladiosum sp. nov. (Japan: Honshu: Fukui Pref.). Additional records of six species are reported from China and Japan. The genus is now represented in the Old World by as many as 620 species, 131 of which have been recorded from Japan. An updated checklist of the Lathrobium species of Japan is provided.

566:
A revision of Palaearctic Lobrathium. VII. A new species from Greece, a revalidation, and additional records (Coleoptera: Staphylinidae: Paederinae). – Lobrathium moreanum sp. nov. (Greece: Pelopónnisos: Menalo Oros), a close relative of L. multipunctum (Gravenhorst, 1802), is described and illustrated. Lobrathium gallienii (Fagniez, 1917), previously a synonym of L. multipunctum, is revalidated. Additional records of seven species from Turkey, Iran, and China are reported. The genus is now represented in the Palaearctic region by 153 species.

568:
Monograph of the Staphylinidae of Crete (Greece). Part I. Diversity and endemism (Insecta:Coleoptera). – With a geological history of long isolation, fragmentation, partial submersion, and subsequent rising, characterized by a remarkable diversity in topology, geology, and habitats, and the fifth-largest island of the Mediterranean, Crete has provided an ideal setting for speciation and developing a unique and diverse Staphylinidae fauna. Although beginning already in the middle of the 19th century, the exploration of this fauna gained momentum only recently. Based on a critical revision of literature data, on material collected during several recent field trips, and on additional previously unpublished records, a checklist of the Staphylinidae of Crete is compiled, including as many as 397 named species, with 115 of them reported from the island for the first time and with 48 new species described in the second part of the monograph. Additional unidentified and unnamed species, as well as species doubtfully or erroneously recorded from Crete are listed separately. The systematic, zoogeographic, and ecological composition of the Staphylinidae fauna is characterized with a special focus on the endemics. The latter are represented by 111 named and at least ten unnamed species, account for nearly one-third of the fauna, and mainly belong to the subfamilies Aleocharinae, Scydmaeninae, Pselaphinae, Paederinae, Leptotyphlinae, Staphylininae, and Omaliinae. More than half of them are even locally endemic. The genera with the greatest number of endemic species are Cephennium Müller & Kunze, 1822 of the Scydmaeninae (twelve named species) and Geostiba Thomson, 1858 of the Aleocharinae (nine named species). More than two-thirds of the endemic species are epigeic inhabitants of forests, bush and shrub habitats, grassland, and wetlands, approximately one-fourth is confined to deeper soil strata or endogean, and 8 % are myrmephilous. Slightly more than half (59 species; 53 %) of the endemics are most likely the result of in-situ radiation (18 lineages), the remainder of vicariance with mainland species. A comparison with other Mediterranean islands revealed that (a) total diversity of the Cretan fauna is significantly greater than that of other East Mediterranean islands except Corfu, but much lower than that of Corsica, Sardinia, and Sicily, (b) regarding the number of endemic species Crete ranks third behind Corsica and Sardinia, and (c) the rate of endemism is greater in Crete than in any other Mediterranean island. Explanations for the observed differences in the diversity and composition of the faunas of the islands are discussed. Stichoglossa graeca Bernhauer, 1905, a species of which previously only the lectotype was known, is redescribed and illustrated. Six synonymies are proposed and one name is revalidated: Phloeocharis longipennis Fauvel, 1875 = P. hummleri Bernhauer, 1915, syn. nov.; Phytosus balticus Kraatz, 1859 = P. holtzi Bernhauer, 1935, syn. nov.; Atheta nigra (Kraatz, 1856) = A. biroi Scheerpeltz, 1964; syn. nov.; Domene stilicina (Erichson, 1840) = D. lohseiana Bordoni, 1977, syn. nov.; Oedichirus rubronotatus Pic, 1903, revalidated = O. reitteri Bernhauer, 1908, syn. nov.; Pseudolathra quadricollis (Fauvel, 1875) = P. cretensis Bordoni, 1986, syn. nov. A lectotype is designated for Stichoglossa graeca Bernhauer, 1905. Two species are reported from Greece for the first time.

569:
Monograph of the Staphylinidae of Crete (Greece). Part II. Descriptions of new species (Insecta:Coleoptera). – One genus, Cretotyphlus Assing gen. nov., and 48 species of Staphylinidae from the Greek island Crete, 47 of them island endemics or even locally endemic, are described and illustrated: Amischa cretica Assing spec. nov., Atheta (Anopleta) digitalis Assing spec. nov., Hydrosmecta insularum Assing spec. nov. (Crete, Ikaría, Lesbos, Samothraki), Geostiba (Sipalotricha) inexsecta Assing spec. nov. (region to the east-southeast of Rethimno), Cousya candica Assing spec. nov. (East Crete: Dikti), Oxypoda (Bessopora) bimontium Assing spec. nov. (Dikti, Psiloritis), O. (Mycetodrepa) retunsa Assing spec. nov., and Typhlocyptus creticus Assing spec. nov. of the Aleocharinae; Geomitopsis cretica Assing spec. nov. (East Crete) of the Osoriinae; Allotyphlus (Moreotyphlus) foedatus Assing spec. nov. (West Crete), Kenotyphlus virgatus Assing spec. nov. (West Crete), Cretotyphlus hamatus Assing spec. nov. (East Crete: Dikti), C. chanianus Assing spec. nov. (region to the southwest of Chania), and C. idanus Assing spec. nov. (Central Crete: Psiloritis) of the Leptotyphlinae; Pseudobium creticum Assing spec. nov. of the Paederinae; Gabrius candicus Assing spec. nov. of the Staphylininae; Cephennium arcuatum Assing spec. nov. (East Crete), C. thripticum Assing spec. nov. (East Crete: Thripti), C. selinonum Assing spec. nov. (Southwest Crete), C. meybohmi Assing spec. nov. (West Crete), C. idanum Assing spec. nov. (Central Crete: Psiloritis), C. sinuosum Assing spec. nov. (East Crete), C. fortespinosum Assing spec. nov. (East Crete), C. hamulatum Assing spec. nov. (East Crete: Dikti), C. curvatum Assing spec. nov. (East Crete), C. selenanum Assing spec. nov. (East Crete: Selena Oros), C. latius Assing spec. nov. (Central Crete: Psiloritis), C. chanianum Assing spec. nov. (West Crete), Euconnus (Tetramelus) zakrius Meybohm spec. nov. (East Crete), Stenichnus (Stenichnus) brachati Meybohm spec. nov., S. (S.) orientalis Meybohm spec. nov. (East Crete), S. (S.) aegialioides Meybohm spec. nov., Leptomastax cretica Meybohm spec. nov., and L. thriptica Meybohm spec. nov. (East Crete: Thripti) of the Scydmaeninae; Afropselaphus doernfeldae Brachat spec. nov. (West Crete: Lefka Ori), A. assingi Brachat spec. nov. (West Crete: Lefka Ori), A. diktianus Brachat spec. nov. (East Crete: Dikti), A. thripticus Brachat spec. nov. (East Crete: Thripti), Amauronyx chanianus Brachat spec. nov. (West Crete), A. occidens Brachat spec. nov. (West Crete), A. askifouicus Brachat spec. nov. (West Crete), Bryaxis meybohmianus Brachat spec. nov. (West Crete), Bythinus creticus Brachat spec. nov. (Central Crete: Psiloritis), Euplectus assingi Brachat spec. nov. (Crete), Faronus meybohmi Brachat spec. nov. (West Crete: Lefka Ori), F. lefkamontium Brachat spec. nov. (West Crete: Lefka Ori), F. albimontium Brachat spec. nov. (West Crete: Lefka Ori), and Tychus chanianus Brachat spec. nov. (West Crete) of the Pselaphinae; Sepedophilus creticus Schülke spec. nov. of the Tachyporinae. Keys to the species of Cephennium Müller & Kunze, 1822 and Stenichnus Thomson, 1859 of Crete are provided. A lectotype is designated for Amauronyx paganettii Blattný & Blattný, 1916.

570:
On the taxonomy and zoogeography of some West Palaearctic Cypha species (Coleoptera: Staphylinidae: Aleocharinae). – Three species are (re-)described and illustrated: Cypha squamipennis (Fauvel, 1902), C. carinthiaca (Scheerpeltz, 1958), and C. bisinuata sp. nov. (South Italy). Based on an examination of types and additional material, C. hanseni (Palm, 1949) is removed from synonymy with C. seminulum (Erichson, 1839) and synonymized with C. pulicaria (Erichson, 1839). A revision of material of species of the newly established C. spathulata group (including C. spathulata Assing, 2007, C. graeca Assing, 2004, C. lindbergi (Palm, 1935), and C. bisinuata) revealed an allopatric distribution pattern in the Mediterranean and Caucasus regions. The distributions of these species are mapped. The male primary sexual characters of C. seminulum, C. pulicaria, and of the species of the C. spathulata group are illustrated. Cypha seminulum is reported from Portugal and Georgia for the first time.

571:
Revision of the Apimela species of the Palaearctic Region with a focus on the West Palaearctic fauna, and with additional notes on the genus Franzidota (Coleoptera: Staphylinidae: Aleocharinae: Oxypodini). – Based on a revision of types and additional material, the oxypodine genus Apimela Mulsant & Rey, 1874 is represented in the Palaearctic region by 29 described species, seven of them confined to the West Palaearctic including Middle Asia, 21 to the East Palaearctic, and one widespread in the southern East Palaearctic and Oriental regions. Two additional species of which neither type material nor illustrations were available remain of uncertain status. Most of the species are (re-)described and/or illustrated, nine of them for the first time: Apimela mutata sp. nov. (= A. macella auctt.) (widespread in the West Palaearctic region); A. wunderlei sp. nov. (South Italy); A. graeca sp. nov. (Greece); A. kirghisica sp. nov. (Kyrgyzstan); A. ussurica sp. nov. (Russian Far East); A. baculata sp. nov. (China: Yunnan); A. bilobata sp. nov. (China: Zhejiang, Sichuan, Yunnan); A. auriculata sp. nov. (China: Yunnan); A. lamellata sp. nov. (North Vietnam). Two genus-group and four species-group synonymies are proposed: Apimela Mulsant & Rey, 1874 = Parapimela Cameron, 1939, syn. nov., = Himalkompsusa Pace, 2006, syn. nov.; Apimela macella (Erichson, 1839) = A. pallens (Mulsant & Rey, 1852), syn. nov., = A. mulsanti (Ganglbauer, 1895), syn. nov.; A. indica (Cameron, 1939) = A. assamensis (Cameron, 1939), syn. nov.; A. chinensis Pace, 1999 = A. tibetana Pace, 2012, syn. nov. The genus-group synonymies result in the following binomina: Apimela indica (Cameron, 1939), comb. nov.; A. persimilis (Cameron, 1939), comb. nov.; A. nepalicola (Pace, 2006), comb. nov.; A. sinica (Pace, 2012), comb. nov.; A. hartmanni (Pace, 2006), comb. nov.; A. taiwanensis (Pace, 2010), comb. nov.; A. morvani (Pace, 1992), comb. nov. “Apimela” lucidula Pace, 1992 belongs to an oxypodine genus of unknown identity and is excluded from Apimela. For various reasons (type material not accessible or not found; descriptions based on females only), several species remain of doubtful identity and partly uncertain generic assignment. Apimela is represented in the Palaearctic region by two distinct clades distinguished primarily by the male and female sexual characters, the A. macella lineage (seven species) and the A. mutata lineage (22 species), both of them distributed across the Palaearctic region. Most Apimela species inhabit the banks of rivers and streams with substrates composed of coarse sand and fine gravel in mountainous regions. An updated catalogue of the Apimela species of the world and a key to the species of the West Palaearctic region including Middle Asia are provided. The distributions of ten species are mapped. In an appendix, the genus Franzidota Pace, 1982 is moved from Oxypodina to Meoticina, and F. formosana sp. nov. (Taiwan) and F. gilva  sp. nov. (Taiwan) are described and illustrated.

572:
On the taxonomy and zoogeography of Paederus. V. Two new species from Laos and China, a new synonymy, new subgeneric assignments, and new records from the Palaearctic region (Coleoptera: Staphylinidae: Paederinae). – Paederus laoticus sp. nov. (Northwest Laos: Bokeo province) and P. (Harpopaederus) willersi sp. nov. (China: Sichuan) are described and illustrated. A new synonymy is proposed: Paederus riparius (Linnaeus, 1758) = P. melanurus Aragon, 1830, syn. nov. Paederus kambaitiensisis Scheerpeltz, 1965 is assigned to the subgenus Heteropaederus Scheerpeltz, 1957 and P. tamulus Erichson, 1840, previously in Eopaederus Scheerpeltz, 1957, is removed from this subgenus and treated as Paederus incertae sedis. The aedeagus of P. kambaitiensis is illustrated for the first time. New records of nine species are reported, among them a new record from the Palaearctic region, two from China, and two new province records from China. The revised distributions of eight species of the subgenus Harpopaederus Scheerpeltz, 1957 are mapped.

573:
On the Tetrasticta fauna of Laos (Coleoptera: Staphylinidae: Aleocharinae: Aleocharini). – Three species of Tetrasticta Kraatz, 1857 are described and illustrated, all of them from the same locality in Bokeo province, Northwest Laos: Tetrasticta varia spec. nov.; T. crassa spec. nov.; T. puncticeps spec. nov. The habitus and primary sexual characters of T. brevipennis (Bernhauer, 1903) are figured. Aside from the new species, four species are reported from Laos, three of them for the first time. Thus, the currently known Tetrasticta fauna of Laos is composed of seven species. Tetrasticta brevipennis is reported from Malaysia, the Indian province Uttarakhand, and the Chinese province Guangdong, and T. bobbii Zheng & Zhao, 2014 from Myanmar for the first time. In an addendum, Tetrasticta borneana spec. nov. (Malaysia: Sabah) is described and illustrated. The genus now includes a total of 20 species, except for one Aftrotropical species all of them distributed in the southern East Palaearctic and Oriental regions.

574:
A revision of Palaearctic and Oriental Rugilus VII.Two new species from China and Laos, and additional records (Coleoptera: Staphylinidae: Paederinae). – Two species of Rugilus Leach, 1819 are described and illustrated: Rugilus (Rugilus) biapicalis spec. nov. (China: Yunnan); Rugilus (Eurystilicus) laoticus spec. nov. (North Laos: Bokeo province). Additional records of five species of the subgenus Eurystilicus Fagel, 1953 are reported, among them two first records from Vietnam. The genus now includes a total of 255 species worldwide, 104 of which are distributed in the Palaearctic and Oriental regions. As many as 36 species, most of them endemic, have been reported from China.

575:
Three new myrmecophiles of the tribes Sahlbergiini and Aenictoteratini from Laos and Malaysia (Borneo) (Coleoptera: Staphylinidae: Aleocharinae). – Malayloeblius laoticus spec. nov. (Northwest Laos) and M. tensus spec. nov. (Borneo: Sabah) of the Sahlbergiini, and Dentaphila iniqua spec. nov. of the Aenictoteratini (Northwest Laos) are described, illustrated, and distinguished from previously described species of the respective genera. Two additional records of Malayloeblius borneensis Pace, 2014 are reported from Borneo. The host ants of the new species are unknown, but available evidence suggests that these species may be associated with army ants.

576:
On the taxonomy of Anomognathus Solier, 1849: the first records of the genus from Laos and a new synonymy (Coleoptera: Staphylinidae: Aleocharinae). – Anomognathus laoticus sp. nov. (Northwest Laos: Bokeo province) and A. triangularis sp. nov. (Northeast Laos: Houa Phan province), the first representatives of the genus to be recorded from Laos, are described and illustrated. A new synonymy is proposed: Placusa nitida Fauvel, 1900 = Anomognathus ispartaensis Örgel, Avci & Özek, 2019; Placusa nitida is reported from Spain, Turkey, and Israel for the first time. The genus Anomognathus Solier, 1849 currently includes 27 valid species distributed in all major zoogeographic regions, except for the Afrotropical region.

577:
On the taxonomy and zoogeography of Anaulacaspis Ganglbauer, 1895 and Borboropora Kraatz, 1862: new species, new records, and the first records of both genera from Laos (Coleoptera: Staphylinidae: Aleocharinae: Falagriini). – Anaulacaspis spinosa sp. nov. (Laos: Bokeo province), A. aberrans sp. nov. (Laos: Bokeo and Xieng Khouang provinces), and Borboropora laotica sp. nov. (Laos: Bokeo province) are described and illustrated. Additional records of nine previously described species of Anaulacaspis Ganglbauer, 1895 and one of Borboropora Kraatz, 1862 are reported, among them four new country records from the West Palaearctic region and one first record from Laos and the Oriental region. Anaulacaspis is recorded from Laos, Borboropora from Laos and the Oriental region for the first time.

578:
On the taxonomy and zoogeography of Chinecallicerus VIII. New species and records, with the first record of the genus from Laos and from outside China (Coleoptera: Staphylinidae: Aleocharinae: Geostibini). – Four species of Chinecallicerus Assing, 2004 are described and illustrated: Chinecallicerus dispersus sp. nov. (China: West Sichuan, West Yunnan); C. unituberculatus sp. nov. (China: Sichuan: Gongga Shan); C. dabaicus sp. nov. (China: Hubei: Daba Shan); C. laoticus sp. nov. (Northwest Laos: Bokeo province). The previously unknown male sexual characters of C. formidabilis Assing, 2019 and the spermathecae of C. serratus Assing, 2009, C. pinnatus Assing, 2017, and C. laevigatus Assing, 2006 are illustrated for the first time. These species, all of them previously known only from their respective holotypes, are reported from additional localities in Sichuan and Yunnan. Chinecallicerus laoticus represents the first record of the genus from Laos, from outside China, and from the whole of the Oriental region, C. dabaicus the first record from the Chinese province Hubei and the easternmost record of the genus. Enormous intraspecific variation was observed for C. dispersus. Including the new species, the genus now includes 22 species, ten of which have been recorded from the Chinese province Yunnan, ten from Sichuan, one from Qinghai, one from Hubei, and one from Laos. As many as 17 species are known only from their respective type localities, twelve of them are represented exclusively by their respective holotypes. A checklist and a newly designed key to species are provided. The distributions of the genus as a whole and of seven individual species are mapped.

579:
Revision of Amarochara VI. New species and records from the Palaearctic and Oriental Regions (Coleoptera: Staphylinidae: Aleocharinae: Aleocharini).  – Three species of Amarochara Thomson, 1858 are described and illustrated: Amarochara sinuosa sp. nov. (North Laos, North Vietnam); A. bifurcata sp. nov. (Northeast Laos: Houa Phan province); A. spinosula sp. nov. (Northeast Laos: Houa Phan province). Additional records of five previously described species are reported, among them new country and new province records. The genus now includes a total of 50 species.

580:
Revision of the Alevonota species of the Palaearctic region IV. Three new species from Sichuan, China (Coleoptera: Staphylinidae: Aleocharinae: Geostibini). – Three species of Alevonota Thomson, 1858, all of them from West Sichuan, China, are described and illustrated: Alevonota pliciventris sp. nov. (Gongga Shan); A. ambulans sp. nov. (Shalui Shan); A. tenuilobata (Erlang Shan). The genus is now represented in the Palaearctic region by 49 species, 28 of them West Palaearctic and 21 East Palaearctic.

581:
On Orsunius IV. Three new species, new records, and the first record of the genus from the Australian region (Coleoptera: Staphylinidae: Paederinae: Medonina). – Three species of the genus Orsunius Assing, 2011 are described and illustrated: Orsunius tortus sp. nov. (Thailand) and O. frischi sp. nov. (Indonesia: Jawa Barat), both of them belonging to the O. wunderlei group, as well as O. maindai sp. nov. (Indonesia: Papua), the first record of the genus from the Australian region. Additional records of nine previously described species are reported, among them five new country records from Thailand (2), China (1), Hong Kong (1), and Cambodia (1). The genus now includes 24 named extant species distributed in the southern East Palaearctic, the Oriental, and the northern Australian regions.

582:
A new species of Lathrobium from Shikoku, Japan (Coleoptera: Staphylinidae: Paederinae). – Lathrobium sugitatense sp. nov. (Japan: Shikoku: Ehime: Mt. Sugitate) is described, illustrated, and compared with its close relative, L. kamezawai Watanabe, 2005. New figures are provided for L. kamezawai. Based on several conspicuous synapomorphies, L. sugitatense and L. kamezawai are hypothesized to represent adelphotaxa. The Lathrobium fauna of Japan is now composed of 134 species, 24 of which are distributed in Shikoku.

583:
On the Cephennium fauna of Greece (Coleoptera, Staphylinidae, Scydmaeninae). – Thirteen species of Cephennium Müller & Kunze, 1822 from Greece are described and illustrated: C. dilatatum nov.sp. (Kavala: Oros Pangéo), C. olympicum nov.sp. (Pieria: Olympos), C. pellucidum nov.sp. (Fthiotis/Evritania: Oros Timfristos), and C. brachati nov.sp. (Flórina) of the C. perispinctum group; C. aciferum nov.sp. (Pelopónnisos, Thessalia, Etolia-Akarnania, Kefalonia, Corfu), C. sagittatum nov.sp. (Ipiros), and C. linguatum nov.sp. (Levkas) of the C. jonicum group; C. cornutum nov.sp. (Fthiothis: Oros Timphristos) of the C. corruptum group; C. vitsiense nov.sp. (Flórina: Oros Vitsi); C. vellicans nov.sp. (Ioánnina: Oros Gramos); C. catax nov.sp. (Thessalia: Oros Ossa); C. dramanum nov.sp. (Drama); C. fibuliferum nov.sp. (Fthiothis, Pelopónnisos). Cephennium jonicum Holdhaus, 1908 and C. petraeum Apfelbeck, 1911, previously subspecies of C. jonicum, are treated as distinct species. Based on drawings by the late Claude Besuchet, the aedeagi of several Cephennium species described prior to 1910, of which published illustrations had not been available, are figured. New records of two previously described species are reported. Including the newly described taxa, the Cephennium fauna of Greece currently includes 35 named species, twelve from Crete (endemic), one from Karpathos (endemic), one from Samothraki (endemic), one from Levkas, and the remainder from the mainland, the Pelopónnisos, and the Ionian Islands Corfu, Levkas, and Kefalonia. A catalogue of the Cephennium species recorded from Greece is provided.

584:
On the Cephennium fauna of the Iberian Peninsula and the Atlantic Islands (Coleoptera, Staphylinidae, Scydmaeninae). – Based on a study of Iberian and Madeiran material of Cephennium Müller & Kunze, 1822, as well as on unpublished notes compiled by the late Claude Besuchet, 38 named species are recognized in Iberian Peninsula to the south of the Spanish border with France and Andorra, and three in Madeira, the only Atlantic archipelago from where the genus has been recorded. The species are attributed to eight species groups. With three exceptions (male unknown, type material lost, or identity doubtful), illustrations or references to recently published figures are provided for all the species of the Iberian Peninsula and Madeira. Seventeen species are described for the first time: C. constrictum nov.sp. (Spain: Cataluña: Sierra del Cadi) of the C. perispinctum group; C. validum nov.sp. (Northwest Spain, North Portugal), C. conlectior nov.sp. (North Portugal: Serra da Estrela), C. algarvense nov.sp. (Portugal), and C. lompei nov.sp. (Madeira proper) of the C. validum group; C. filabresicum nov.sp. (Spain: Andalucía: Sierra de los Filabres) and C. espunae nov.sp. (Spain: Murcia: Sierra de Espuña) of the C. caecum group; C. tensum nov.sp. (Spain: Alicante: Sierra de Aitana) and C. andujari nov.sp. (Spain: Alicante: Sierra de Aitana) of the C. tensum group; C. tenuissimum nov.sp. (Spain: Murcia: Sierra de Espuña) of the C. tenuissimum group; C. alacantae nov.sp. (Spain: Alicante), C. cujonicum nov.sp. (Spain: Castilla-La Mancha: Sierra de Segura), C. verrens nov.sp. (Spain: Andalucía: Sierra de Segura), C. segurae nov.sp. (Spain: Castilla-La Mancha: Sierra de Segura), C. nevadae nov.sp. (Spain: Andalucía: Sierra Nevada), C. cazorlae nov.sp. (Spain: Andalucía: Sierra de Cazorla), and C. malagae nov.sp. (Spain: Andalucía: Malaga) of the subgenus Macroderus Croissandeau, 1894. Two subspecies of C. pygmaeum Saulcy, 1870 are regarded as doubtful. One species is recorded from Spain for the first time. A catalogue of the named Cephennium species recorded from the Iberian Peninsula and Madeira is provided. The distributions of 23 species are mapped. Including the new taxa, the genus is now represented by 251 named species.

585:
On the taxonomy of Parocyusa, Tectusa, and miscellaneous genera of Oxypodina (Coleoptera: Staphylinidae: Aleocharinae: Oxypodini). – A comparative morphology-based study of types and additional material of species previously assigned to Tetralaucopora Bernhauer, 1928, Tectusa Bernhauer, 1899, and various other genera of Oxypodina results in numerous taxonomic changes. Parocyusa Bernhauer, 1902, previously a synonym of Tectusa, is revalidated. The genus has a Holarctic distribution and currently includes 33 described species, 20 species in the West Palaearctic (including Middle Asia), twelve in the East Palaearctic, and two species (one of them probably native in the West Palaearctic) in the Nearctic region. The validity of two species, one from North Europe and one from Japan, requires confirmation and one name is treated as a tentative synonym. Ten of the West Palaearctic species are micropterous or submacropterous and have restricted distributions, whereas the remainder is macropterous and more or less widespread. Tectusa presently includes 41 described species, all of them micropterous and with restricted distributions, with the vast majority confined to mountain ranges in the Balkans. The genus is most likely polyphyletic. However, with the type material of several species (including the type species of available genus-group names) currently inaccessible, this issue will have to be addressed in a future revision. New combinations are proposed for all the species previously in Tetralaucopora and some previously in Tectusa or other genera, nine names are synonymized, and one name is preserved according to Article 23.9 of the Code (ICZN 1999): Parocyusa Bernhauer, 1902 = Tetralaucopora Bernhauer, 1928, resyn.; Parocyusa antennata (Eppelsheim, 1878), comb.n.; P. baicalensis (Eppelsheim, 1893), comb.n. = beijingensis (Pace, 1999), syn.n., = hebeiensis (Pace, 1999), syn.n.; P. bicolorata (Assing, 2007), comb.n.; P. caligula (Assing, 1996), comb.n.; P. carnica Lohse, 1988; P. championi (Cameron, 1939), comb.n.; P. crebrepunctata (Strand, 1962), comb.n.; P. fuliginosa (Casey, 1906), comb.n.; P. germana (Cameron, 1939), comb.n. = rougemonti (Pace, 1986), syn.n.; P. hartmanni (Pace, 2013), comb.n. (ex Ocalea); P. holdhausi (Bernhauer, 1902); P. japonica (Cameron, 1933), comb.n.; P. knabli (Bernhauer, 1914) = tirolensis Scheerpeltz, 1958, syn.n.; P. lebedevi (Bernhauer, 1928), comb.n. = bucharica (Bernhauer, 1928), syn.n.; P. longicollis (Eppelsheim, 1889), comb.n.; P. longitarsis (Erichson, 1839), comb.n., nomen protectum = attenuata (Stephens, 1832), nomen oblitum, = syriaca (Saulcy, 1865), syn.n.; P. montana (Kraatz, 1856), comb.n.; P. ripicola (Cameron, 1939), comb.n.; P. rubicunda (Erichson, 1837), comb.n. = cingulata (Kraatz, 1856), syn.n.; P. schuelkei (Assing, 1996), comb.n.; P. strupiiana Scheerpeltz, 1958; P. subcyanea (Cameron, 1939), comb.n.; P. yakouensis (Pace, 2010), comb.n. (ex Ocyusa); P. yunnanensis (Pace, 1993), comb.n.; “Ocyusa” fuscobrunnea (Cameron, 1939), comb.n.; “Cousya” quadrisulcata (Bernhauer, 1935) = nepalensis Pace, 2006, syn.n.; Oxypoda sinonigra (Pace, 2017), comb.n.; Oxypoda besucheti Focarile, 1982. Numerous species of Parocyusa and Tectusa are (re-)described and illustrated, among them 19 new taxa (nine of Parocyusa and ten of Tectusa): Parocyusa dilatata sp.n. (China: Yunnan); P. gilvipennis sp.n. (Kazakhstan); P. gonggaica sp.n. (China: Sichuan); P. gracillima sp.n. (South Turkey); P. kahleni sp.n. (South Slovenia); P. maculipennis sp.n. (Northwest Iran); P. matarata sp.n. (South Iran); P. spinosa sp.n. (China: Gansu); P. virilis sp.n. (Caucasus region: Russia, Armenia); Tectusa acrilobata sp.n (Erimanthos); T. apanoica sp.n. (Menalo); T. chelmosica sp.n. (Aroania); T. fimbriata sp.n. (Panahaiko); T. latilobata sp.n. (Giona, Iti, Parnassos); T. menaloica sp.n. (Menalo); T. parnonica sp.n. (Parnon); T. pauli sp.n. (Aroania); T. taygetana sp.n. (Taygetos); T. vodiasa sp.n. (Panahaiko). All the newly described Tectusa species are distributed in Greece and, except for T. latilobata, all of them in the Pelopónnisos. The Carpathian and one Anatolian species previously attributed to Tectusa are moved to the genus Duplocyusa gen.n. (type species Oxypoda uludaghensis Fagel, 1971), with the Carpathian species in the subgenus Carpocyusa subgen.n. (type species Tectusa nigromontis Zerche, 2007): Duplocyusa (Duplocyusa) uludaghensis Fagel, 1971, comb.n.; Duplocyusa (Carpocyusa) nigromontis (Zerche, 2007), comb.n.; D. (C.) bucegiensis (Zerche, 2007), comb.n.; D. (C.) transsylvanica (Zerche, 2007), comb.n.; D. (C.) rosenauensis (Zerche, 2007), comb.n.; D. (C.) ceahlauensis (Zerche, 2007), comb.n.; D. (C.) rodnaensis (Zerche, 2007), comb.n.; D. (C.) calimaniensis (Zerche, 2008), comb.n.; D. (C.) iucasensis (Zerche, 2008), comb.n. In an appendix on miscellaneous genera of Oxypodina, the following revalidation, synonymies, and new combinations are proposed, and five species are described: Eurylophus J. Sahlberg, 1876, revalidated = Drepasiagonusa Pace, 2012, syn.n.; Eurylophus grandiceps J. Sahlberg, 1876, comb.n. = feldmanni (Assing, 2018), syn.n.; Eurylophus smetanai (Pace, 2012), comb.n.; E. procerus (Assing, 2018), comb.n.; E. tibeticus (Assing, 2018), comb.n.; E. angulatus (Assing, 2018), comb.n. (all Eurylophus species except E. grandiceps ex Drepasiagonusa); “Cousya” lobifera Pace, 2015 (probably belonging to an undescribed genus); Cousya luteipennis sp.n. (Uzbekistan); Parocalea tenebricosa sp.n. (Russian Far East); Trichoglossina taiwafimbriata (Pace, 2010), comb.n. (ex Oxypoda); Trichoglossina retunsa sp.n. (China: Yunnan); T. tricuspidata sp.n. (China: Yunnan); Chinecousya globosa sp.n. (China: Yunnan). Lectotypes are designated for Chilopora antennata Eppelsheim, 1878, C. baicalensis Eppelsheim, 1893, C. ripicola Cameron, 1939, and C. championi Cameron, 1939. The distributions of Parocyusa rubicunda, a species with evidently parthenogenetic populations in large parts of its range, and of 17 species of Tectusa are mapped. Numerous new records are reported.

586:
On the Lomechusini of the Palaearctic and Oriental regions. XXVIII. New genera, species, and records, with a focus on the fauna of Laos and Borneo (Coleoptera: Staphylinidae: Aleocharinae). – Based on a morphological study of material of 14 genera of Lomechusini from the Oriental and East Palaearctic regions, primarily from Laos and Borneo, and of some specimens from Irian Jaya, numerous species are described and illustrated, 78 of them for the first time: Orphnebius abbreviatus sp. nov. (Laos), O. crassisetosus sp. nov. (Laos), O. taleatus sp. nov. (Borneo), O. geminus sp. nov. (Borneo), O. sextuberculatus sp. nov. (Borneo), O. retornatus sp. nov. (Borneo), O. derectus sp. nov. (Borneo), O. comatus sp. nov. (Laos), O. calvus sp. nov. (Laos), O. baculifer sp. nov. (Borneo), O. unguicus sp. nov. (Borneo), O. curvatus sp. nov. (Borneo), O. varicornutus sp. nov. (Laos), O. biacer sp. nov. (Borneo), O. conifer sp. nov. (Borneo), O. mollis sp. nov. (Borneo), O. linguis sp. nov. (Borneo), O. sinulinguis sp. nov. (Borneo), O. cuspilinguis sp. nov., O. colitus sp. nov. (Borneo), O. tensus sp. nov. (Borneo), and O. pressatus sp. nov. (Borneo) of the Orphnebius hauseri group; Orphnebius (Deroleptus) transfigens sp. nov. (Borneo), O. (D.) aciformis sp. nov. (Borneo), O. (D.) acifer sp. nov. (Borneo), O. (D.) quadricultratus sp. nov. (Borneo), O. (D.) floreni sp. nov. (Borneo), O. (D.) obelifer sp. nov. (Borneo), O. (D.) nodatus sp. nov. (Borneo), O. (D.) filiformis sp. nov. (Borneo), O. (D.) plexus sp. nov. (Borneo), O. (D.) tuberculifer sp. nov. (Borneo), O. (D.) tuberosus sp. nov. (Borneo), O. (D.) granulosus sp. nov. (Borneo), O. (D.) verrucosus sp. nov. (Borneo), O. (D.) botulus sp. nov. (Borneo), O. (D.) pinnatus sp. nov. (Borneo), O. (D.) bicarinatus sp. nov. (Borneo), O. (D.) immutatus sp. nov. (Laos), O. (D.) buccatus sp. nov. (Laos), O. (D.) innexus sp. nov. (Borneo), O. (D.) furviceps sp. nov. (Borneo), O. (D.) compressus sp. nov. (Borneo), O. (D.) bituberculatus sp. nov. (Borneo), O. (D.) tensicornis sp. nov. (Borneo), O. (D.) rogans sp. nov. (Borneo), O. (D.) tuberifer sp. nov. (Borneo), O. (D.) bulbifer sp. nov. (Sulawesi Tengah), O. (D.) bulbosus sp. nov. (Philippines: Mindanao), and O. (D.) hirsutus sp. nov. (Borneo); Amaurodera dentissima sp. nov. (Myanmar), A. setosicollis sp. nov. (Borneo), and A. floreni sp. nov. (Borneo); Drusilla trituberculata sp. nov. (China: Hubei), D. longispinosa sp. nov. (Borneo), D. iniqua sp. nov. (Borneo), and D. samalica sp. nov. (Philippines: Mindanao); Witteia arboris sp. nov. (Borneo); Tensusa procera sp. nov. (Laos); Pheidologitonetes quadriplicatus sp. nov. (Laos), P. artiplicatus sp. nov. (Laos), P. acer sp. nov. (Vietnam), P. retunsus sp. nov. (Vietnam), P. sagittatus sp. nov. (Laos), and P. punctatus sp. nov. (Laos); Zyras (Zyras) bituberosus sp. nov. (Laos), Z. (Z.) semicalvus sp. nov. (Laos), Z. (Z.) janetlarae sp. nov. (Borneo), Z. (Z.) parvihirtus sp. nov. (Borneo), Z. (Z.) cyaniceps sp. nov. (Irian Jaya), Z. (Z.) maindai sp. nov. (Irian Jaya), Z. (Z.) interruptus sp. nov. (Irian Jaya), Z. (incertae sedis) pressitibialis sp. nov. (Borneo), and Z. (i. s.) cuneatus sp. nov. (Laos); Mimaenictus bisetosus sp. nov. (Thailand); Scapaenictus longiscapus sp. nov. (Laos); Planusa granulosa sp. nov. (Laos); Sulciusa boops sp. nov. (Borneo). Four monotypical genera are described: Tensusa gen. nov. (type species: T. procera); Scapaenictus gen. nov. (type species: S. longiscapus); Planusa gen. nov. (type species: P. granulosa); Sulciusa gen. nov. (type species: S. boops). Nine new combinations are established: Orphnebius mirabilis (Pace, 2014), comb. nov. (ex Keratodegnathus Pace, 2014); Pheidologitonetes thaifuscicollis (Pace, 2012), comb. nov. (ex Drusilla Leach, 1819); P. rubricollis (Cameron, 1939), comb. nov. (ex Episkiodrusilla Pace, 2013); P. malayrubricollis (Pace, 2013), comb. nov. (ex Episkiodrusilla); P. neocoenonicacollis (Pace, 2008), comb. nov. (ex Episkiodrusilla); P. vedda Pace, 1992, comb. nov. (ex Drusilla); P. veluticollis (Pace, 2013) , comb. nov. (ex Episkiodrusilla); Drusilla nilgiriensis (Pace, 2001), comb. nov. (ex Pheidologitonetes); D. bartolozzii (Pace, 2001), comb. nov. (ex Pheidologitonetes). Four new species groups are proposed within Orphnebius Motschulsky, 1858: the O. mollis, O. colitus, and O. tensus subgroups of the O. hauseri group, and the O. hirsutus species group of Deroleptus Bernhauer, 1915. The intrageneric affiliations of Orphnebius, their systematic implications, and their nomenclatural consequences are discussed. Two synonymies are established: Deroleptus Bernhauer, 1915 = Aulacothoracobius Bernhauer, 1929, syn. nov.; Pheidologitonetes Cameron, 1939 = Episkiodrusilla Pace, 2013, syn. nov. Updated catalogues of Orphnebius sensu lato and of Pheidologitonetes are provided. Additional records of 59 previously described species are reported, among them numerous new country records.