For corresponding references and
abstracts nos. 1 - 300 see publications ,
abstracts
I, abstracts II, abstracts
III, abstracts IV,
abstracts V, abstracts VI, and
abstracts VII, respectively.
408:
A
new species of Dysanabatium Bernhauer and additional records of D.
jacobsoni Bernhauer
(Coleoptera, Staphylinidae, Paederinae).
Dysanabatium hainanense Peng
& Li, sp. n. (Hainan: Wuzhi Shan, Diaoluo Shan) is described and
illustrated. Additional records of D. jacobsoni Bernhauer, 1915
are reported. The habitus, the sexual characters, and the distribution of this
enormously variable species are illustrated.
409:
On
the Staphylinidae of Sardinia, Italy (Insecta: Coleoptera).
Recently
collected material from Sardinia, comprising more than 1200 specimens and more
than 70 species, is studied. Two species are described and illustrated: Astenus
(Eurysunius)
wunderlei nov.sp. and Leptusa (Adexiopisalia)
meybohmi nov.sp. One synonymy is proposed: Anthobium atrocephalum
(Gyllenhal
1827) = A. sardoum (Scheerpeltz
1961) nov.syn. Records of zoogeographic interest are reported for 19 species,
among them two new records for Sardinia. The currently known distribution of Oxypoda
islandica Kraatz
1857 is mapped.
410:
A
revision of Palaearctic Lobrathium. V. Three new species from the
Himalaya and China, a new synonymy, and additional records (Coleoptera:
Staphylinidae: Paederinae).
Four
new species of Lobrathium Mulsant
& Rey 1878 are described and illustrated: L.
subcaeruleum nov.sp. (North India, Nepal), L.
wutaium nov.sp. (China: Shanxi: Wutai Shan), L.
flexum nov.sp. (China: Jiangxi: Jinggang Shan), and L. circumclusum nov.sp. (Myanmar). Lobrathium zonale Li,
Solodovnikov & Zhou 2013, nov.syn. is placed in synonymy with L.
configens Assing
2012. Additional records of thirteen species are presented, among them the first
record of the genus from Thailand. Lobrathium is now represented in the
Palaearctic region by 135 species and one subspecies, with 21 species recorded
from the Himalaya and 40 from mainland China.
411:
A
revision of Trisunius IV. Two new species and additional records
from China
(Coleoptera: Staphylinidae: Paederinae).
Two
species of the medonine genus Trisunius
Assing
2011 from China are described and illustrated: Trisunius
smetanai nov.sp. (Yunnan: Gaoligong Shan) and T.
rastratus nov.sp. (Hubei: Daba Shan). Additional records of three species
are reported from Yunnan. The East Palaearctic genus now includes 19 species.
412:
A
revision of Palaearctic and Oriental Rugilus. IV. Three new species from
Nepal and additional records (Coleoptera: Staphylinidae: Paederinae).
Three
species of the paederine genus Rugilus Leach
1819 from Nepal are described and illustrated: R. (Rugilus) bibarbatus
nov.sp. (W-Nepal: Karnali and Seti provinces); R. (R.) schmidti
nov.sp. (Manaslu range); R. (R.) bisinuosus nov.sp.
(Manaslu range). Additional records of 17 previously described species are
reported from the Palaearctic and Oriental regions, among them three new country
records. Including the new species, Rugilus is now represented in the
Palaearctic and Oriental regions by 97 species and one subspecies.
413:
On
Orsunius II. Eight new species and additional records (Coleoptera:
Staphylinidae: Paederinae: Medonina).
Eight
species of the Oriental medonine genus Orsunius Assing 2011 are described and illustrated: O. spathulatus
nov.sp. (Philippines: Mindanao); O. stimulans nov.sp. (North Vietnam,
Thailand); O. latexcisus nov.sp. (South Thailand); O. gladiator
nov.sp. (Thailand); O. flavovirgatus nov.sp. (South Thailand); O.
granulosus nov.sp. (Laos, Thailand, North Vietnam); O. cuneus nov.sp.
(Laos); O. heissi nov.sp. (Malaysia: Kedah). Additional records of two
previously named species are reported, among them the first records of O.
immsi (Bernhauer 1914) from
Laos and Thailand. The distributions of all the species of the genus are mapped.
A new key to species and an updated catalogue are provided.
414:
Seven
new species and additional records of Stilicoderus and Stiliderus
(Coleoptera: Staphylinidae: Paederinae).
Five species of Stilicoderus Sharp
1889 and two of Stiliderus Motschulsky
1858 are described and illustrated: Stilicoderus dilatatus nov.sp.
(Taiwan) of the S. japonicus group; Stilicoderus seticollis
nov.sp. (Indonesia: Maluku: Morotai), S. sociabilis nov.sp. (Maluku:
Morotai), S. brevisetosus nov.sp. (Maluku: Morotai), and S.
transversus nov.sp. (Maluku: Halmahera) of the S. hieroglyphicus
group; Stiliderus aviformis nov.sp. (NW-Thailand) of the S.
cicatricosus group; S. tridentatus nov.sp. (Indonesia: Sulawesi
Utara) of the S. brendelli group. Additional records of ten described
species of Stilicoderus and of four species of Stiliderus are
reported.
415:
On
Domene scabripennis ROUGEMONT
and
its close relatives (Coleoptera: Staphylinidae: Paederinae).
Five species of Domene Fauvel
1873, tentatively attributed to the subgenus Macromene Coiffait
1982, from the East Palaearctic and the northern Oriental regions are (re-)
described and illustrated: D. scabripennis Rougemont
1995 (Taiwan), D. alesiana nov.sp. (Taiwan), D. immarginata
nov.sp. (China: Yunnan), D. sagittata nov.sp. (North Vietnam), and D.
firmicornis nov.sp. (China: Zhejiang). These species are assigned to the
newly established D. scabripennis group. The placement of D.
firmicornis in this group, however, is doubtful and requires confirmation. A
key to species is provided. Their distributions are mapped.
416:
Three
new species and a new species group of Medon
from
China and Taiwan (Coleoptera: Staphylinidae: Paederinae).
Three species of the newly established Medon profundus group are
described and illustrated: M. profundus nov.sp. (China: border region
between Shaanxi and Chongqing, Sichuan), M. yunnanicus nov.sp. (China:
Yunnan), and M. alesi (Taiwan). They are evidently closely related and
undoubtedly form a monophyletic group. Their known distributions are mapped. The
preoccupied name Medon flavomarginatus Assing
2013 is replaced with M. gilvimarginatus nov.nom. Medon Stephens
1833 is now represented in the Palaearctic region by 81 confirmed species.
417:
A
Crataraea species associated with Formica chinensis (Coleoptera:
Staphylinidae: Aleocharinae).
Crataraea
myrmecophila
nov.sp. (China: Qinghai) is described and illustrated. The
species was exclusively collected from nests of the ant Formica chinensis
Wheeler 1913. It represents the
first record of pronounced myrmecophily in the genus and the first record of the
genus from the East Palaearctic region east of Middle Asia. Crataraea Thomson
1858 is now represented in the Palaearctic region by five species.
418:
New
species and records of Lathrobium from the Palaearctic region, primarily
from Nepal (Coleoptera: Staphylinidae: Paederinae).
Eighteen
species of Lathrobium Gravenhorst,
1802 from Nepal (16 species) and China (2 species) are described and
illustrated: L. semicirculatum sp. n. (E-Nepal; L. nepalense
group); L. constrictum sp. n. (E-Nepal; L. discissum group); L.
descendens sp. n. (W-Nepal; L. deuvei group); L. ascendens sp.
n. (W-Nepal; L. deuvei group); L. rude sp. n. (C-Nepal:
Dhaulagiri; L. emodense group); L. latibasale sp. n. (W-Nepal; L.
pectinatum group); L. eques sp. n. (W-Nepal; L. pectinatum
group); L. equitans sp. n. (C-Nepal: Dhaulagiri; L. pectinatum
group); L. abruptum sp. n. (W-Nepal; L. pectinatum group); L.
kemum sp. n. (C-Nepal: Dhaulagiri; L. pectinatum group); L.
flexaculeatum sp. n. (C-Nepal: Dhaulagiri; L. pectinatum group); L.
rectaculeatum sp. n. (C-Nepal: Dhaulagiri; L. pectinatum group); L.
curvaculeatum sp. n. (C-Nepal: Dhaulagiri; L. pectinatum group); L.
hebes sp. n. (C-Nepal: Dhaulagiri; L. pectinatum group); L.
imberbe sp. n. (W-Nepal; L. pectinatum group); L. plicatum sp.
n. (C-Nepal: Dhaulagiri; L. pectinatum group); L. glabrimpressum
sp. n. (China: Yunnan: Gaoligong Shan; group assignment uncertain); L.
smetanai sp. n. (China: Gansu: Dalija Shan; L. lentum group). Based
on new material, a redescription and new illustrations of L. aculeatum Coiffait,
1982 are provided. Recently collected material of L. calcaratum Assing,
2012 suggests that the holotype of this species may have been mislabelled and
that, consequently, the type locality is erroneous. The species previously
assigned to the L. aculeatum group are moved to the L. pectinatum
group, and the L. aculeatum group is discarded. Several species of the L.
pectinatum group were found to be subject to a sexual dimorphism of body
size. Additional records of six named species are reported from the West
Caucasus (2 species) and from Nepal (4 species). Including the newly described
species, the genus is currently represented in the Himalaya by 68 species in ten
species groups, in China by 189 species, and in the Palaearctic region by 536
named species and ten subspecies.
419:
A
revision of the species
of Pinobius MacLeay,
1871 of the Oriental, Palaearctic, and Australian Regions (Coleoptera:
Staphylinidae: Paederinae).
The
species of the dolicaonine genus Pinobius MacLeay, 1871 (Coleoptera: Staphylinidae: Paederinae:
Dolicaonina) of the Oriental, Palaearctic, and Australian Regions are revised.
In all, 28 species are recognized, among them 14 species new to science: P.
baculatus sp.n. (Indonesia: Java, Sumba), P. bifidus sp.n. (North
India), P. bihamatus sp.n. (Myanmar), P. brevincisus sp.n. (South
Vietnam), P. carinatus sp.n. (India: Maharashtra), P. discrepans
sp.n. (Australia), P. extensus sp.n. (Papua New Guinea), P.
firmilobatus sp.n. (Thailand, Singapore, Peninsular Malaysia, Sabah), P.
formidabilis sp.n. (Malaysia), P. imberbis sp.n. (India: Assam), P.
inaequalis sp.n. (Indonesia: Sulawesi), P. linguatus sp.n.
(Bangladesh), P. rotundatus sp.n. (India: Assam), P. subaequalis
sp.n. (Indonesia: Sulawesi). All the species are (re-)described and illustrated.
Pinobius vicinus (Kraatz,
1859), previously a synonym of P. indicus (Kraatz,
1859), is revalidated. The following synonymies are proposed: P. indicus
(Kraatz, 1859) = longipennis
(Cameron, 1931), syn.n.; P.
paricolor (Fauvel, 1878) = similis
Last, 1984, syn.n.; P.
sparsiventris (Fauvel, 1886) = javanus
(Cameron, 1936), syn.n. Pinobius
pedatus (Lea, 1923) is
tentatively regarded as a synonym of P. alatus (Lea,
1923). Lectotypes are designated for Dolicaon adjacens Cameron,
1937, D. alatus Lea, 1923, D.
indicus Kraatz, 1859, D.
javanus Cameron, 1936, D.
robustus Kraatz, 1859, D.
sparsiventris Fauvel, 1886, and
D. vicinus Kraatz, 1859. The
revision revealed that the previously described species had largely been
misinterpreted, suggesting that nearly all literature records other than
original descriptions are based on misidentifications. Primarily based on the
morphology of the aedeagus, the species are assigned to four species groups. The
zoogeography of the genus in the study region is discussed. The Indian
Subcontinent, eastwards to Myanmar, is the region with the greatest diversity.
The distributions of all the species are mapped. The examined material was
largely collected at light sources, partly also sifted from leaf litter, at low
to intermediate elevations. Only one of the studied specimens is teneral. A key
to species and a checklist of the Pinobius species of the Oriental,
Palaearctic, and Australian Regions are provided. The genus now comprises a
total of 35 species, ten in the Afrotropical, twenty in the Oriental, and seven
in the Australian Region. Only five species, none of them exclusive, have been
recorded from the south of the Palaearctic Region (southern Arabian Peninsula,
South Himalaya, South China), one of them of Afrotropical and the other four of
Oriental affiliations. In an appendix, a dolicaonine species from the
Afrotropical Region is described: Jarrigeus verberans sp.n. (Nigeria).
420:
A
revision of Nazeris. VI. On the fauna of East Yunnan, China (Coleoptera:
Staphylinidae: Paederinae).
A
recent field trip to eastern Yunnan (Southwest China), a region from where only
one species of Nazeris Fauvel,
1873 had been known, yielded thirteen undescribed Nazeris species, eleven
of which are described and illustrated: N. brevilobatus sp. n. (Dongchuan
env.), N. virilis sp. n. (Wuding env.), N. constrictus sp. n.
(Gejiu env.), N. claviger sp. n. (Gejiu env.), and N. discissus
sp. n. (Pingbian env.) of the newly established N. brevilobatus group; N.
lamellatus sp. n. (Pingbian env.) of the newly established N. lamellatus
group; N. semifissus sp. n. (Pingbian env.) of the newly established N.
semifissus group; N. fibulatus sp. n. (Kunming env.), N. bulbosus
sp. n. (Kunming env.), N. clavilobatus sp. n. (Dongchuan env.), and N.
conicus sp. n. (Xundian env.) of the N. cangicus group. Together with
N. zhangi Watanabe & Xiao,
1993, the latter three species form a distinct lineage within the N. cangicus
group. Two species remain unnamed, as they are represented exclusively by
females. The general distribution of the genus in Yunnan and the individual
distributions of the newly described species are mapped. Nazeris now
includes a total of described 233 species (plus seven subspecies), 118 of which
have been recorded from mainland China. The fauna of Yunnan currently comprises
45 described species.
421:
Three
new species and additional records of Echiaster from Asia (Coleoptera:
Staphylinidae: Paederinae).
Three species of the paederine genus Echiaster
Erichson, 1839 from Asia are
described and illustrated: Echiaster
bivirgatus nov.sp. (Thailand, North India), E. paulexcisus nov.sp.
(Thailand), and E. discrepans nov.sp. (South India). Additional records
from Thailand, Taiwan, and China are reported for E. unicolor Bernhauer,
1922 and E. maior Assing,
2013. The genus is now represented in Asia by five species in two lineages, one
of them composed of one species (E. discrepans) from South India and the
other of the remaining four species. A key to species is provided. The
distribution of the genus in Asia is mapped.
422:
A
revision of Neosclerus Cameron
IV. Two new species and additional records from Taiwan (Coleoptera:
Staphylinidae: Paederinae).
Two micropterous species of Neosclerus
Cameron,
1924 from Taiwan (Nantou Hsien) are described and illustrated, N.
brevispinosus
nov.sp. and N.
biaculeatus
nov.sp. Additional records of N.
bifidus
Assing,
2011 are reported from Taiwan. Including the new species, the
genus now includes 31 species and is represented in Taiwan by nine locally
endemic species.
423:
Two
new species of Eccoptoglossa from Iran and Kyrgyzstan (Coleoptera:
Staphylinidae: Aleocharinae).
Eccoptoglossa
amischoides
nov.sp. (Iran: Ardabil, Lorestan) and E. kirgisica nov.sp. (Kyrgyzstan:
Osh) are described and illustrated. Including the new species, the Palaearctic
falagriine genus Eccoptoglossa Luze,
1904 now comprises five species and is distributed from the East Caucasus region
across Middle Asia eastwards to northwestern Mongolia. The distributions of the Eccoptoglossa
species are mapped and a key to species is provided.
424:
On
the genus Crataraea (Coleoptera: Staphylinidae: Aleocharinae: Oxypodini).
Three species previously attributed to the Crataraea Thomson, 1858, subtribe Microglottina, are revised and
illustrated. Only one of them, C. solskyi (Eppelsheim, 1888), is retained in the genus. Two species are
moved to Aleochara Gravenhorst
1802: Aleochara (Xenochara) rubripennis (Fauvel,
1872), nov.comb., and A. (X.) eremita (Peyerimhoff,
1931), nov.comb. The previous synonym Aleochara rubripes Blatchley,
1910 is revalidated to replace the secondary junior homonym Aleochara
rubripennis (Casey, 1906). A
lectotype is designated for Crataraea rubripennis Fauvel,
1872. Crataraea now includes three species native in the Palaearctic
region, with one species adventive in North America. A key to species is
provided.
425:
A
revision of Tetartopeus IV. A new species from Turkey, new synonymies,
and additional records (Coleoptera: Staphylinidae: Paederinae).
Material of the paederine genus Tetartopeus
Czwalina, 1888 from the
Palaearctic, Oriental, and Australian regions is examined. Tetartopeus korgei nov.sp.
(Turkey: Kars), T. lomnickii (Roubal,
1913), and the previously unknown female secondary sexual characters of T.
lentus Ryvkin, 1989 are (re-)described and illustrated. The
following synonymies are established: Tetartopeus gracilentus (Kraatz,
1859) = Lathrobium pallipes Sharp,
1889, nov.syn., = L. maculatum Last,
1984, nov.syn., = Lobrathium wui Zheng, 2001, nov.syn. = L.
bimaculatum Li, Tang & Zhu, 2007,
nov.syn. The remarkably vast distribution of T. gracilentus, a species
present in the Oriental (Sri Lanka), East Palaearctic, and Australian regions
(New Guinea), is mapped. Additional records of fourteen species are presented,
among them a new country record.
426:
On
the Staphylinidae of Turkey X. Two new species and additional records (Insecta:
Coleoptera).
Recently collected material from Turkey, comprising nearly 700 specimens and
approximately 130 species, as well as older material from the Korge collection
(Berlin), which is now publicly accessible, is studied. Two species of
Paederinae and Staphylininae are described and illustrated: Lobrathium
unidentatum nov.sp. (Turkey: Hakkβri) and Atanygnathus pullus
nov.sp. (Turkey: Adana; Greece: Pelopσnnisos); Israel). Four species are
reported from Turkey for the first time. The distributions of the newly
described species are mapped.
427:
A
new species of Platydomene from North India (Coleoptera: Staphylinidae:
Paederinae).
Platydomene
shavrini
nov.sp. from North India (Uttarakhand), the first Platydomene species to
be recorded from the Himalaya, is described and illustrated. The genus Platydomene
Ganglbauer, 1895 is now represented
in the Palaearctic region by 27 species and three subspecies, of which fourteen
species are distributed in the East Palaearctic.
428:
A
revision of Palaearctic and Oriental Pseudolathra. IV. New species, new
combinations and additional records (Coleoptera: Staphylinidae: Paederinae).
Type and non-type material of Pseudolathra Casey,
1905 from the Palaearctic, Oriental, and Australian regions is revised. Four
species from Papua New Guinea are transferred from Lathrobium Gravenhorst,
1802 to Pseudolathra, redescribed, and illustrated: Pseudolathra
sepikensis (Last, 1984),
nov.comb., P. naviculum (Last,
1984), nov.comb, P. puncta (Last,
1984), nov.comb., and P. cyanea (Last,
1984), nov.comb. Two species are described and illustrated: P. armata
nov.sp. (Peninsular Malaysia) and P. bisinuata nov.sp. (Malaysia:
Borneo). Additional records of five species are reported.
429:
Three
new species, redescriptions, a new synonymy, and additional records of Cypha
from the West Palaearctic region (Coleoptera: Staphylinidae: Aleocharinae).
Five species of Cypha Leach,
1819 from the West Palaearctic region are (re-)described and illustrated: Cypha
rubripennis (Pandellι, 1869)
(France, Spain, Morocco); C. rubicunda (Reitter,
1887) (Turkmenistan); C. truncata nov.sp. (Ukraine); C. prominens
nov.sp. (Tunisia); C. persica nov.sp. (Iran). Cypha tbilisiensis Dauphin,
2005 nov.syn. is placed in synonymy with C. tarsalis (Luze,
1902). Several new country records are reported. The genus is now represented in
the Palaearctic region by 50 described species.
430:
On
the Staphylinidae of Israel II, with a revision of some species of Dinusa
Saulcy
(Coleoptera: Staphylinidae: Aleocharinae).
The results of extensive pitfall trap studies conducted in Israel in 2010 and
2011 are presented, and material of some species of Dinusa Saulcy,
1865 is revised. Records of zoogeographic interest of 49 species are reported,
among them 25 first records from Israel, three from Jordan, three from Iraq, one
from Syria, one from Egypt, and one from Malta. Three species of Paederinae and
Aleocharinae from Israel and one species of Aleocharinae from southern Turkey
are described and illustrated: Astenus (Eurysunius) affimbriatus
nov.sp. (Israel: Hula valley), Pseudosemiris torta nov.sp. (Israel:
numerous localities), Geostiba (Sibiota) meronica nov.sp.
(Israel: Mount Meron), and Dinusa subangulata nov.sp. (Turkey:
Gaziantep). The distributions of Astenus affimbriatus and Pseudosemiris
torta are mapped. Astenus (Eurysunius) platynotus (Saulcy,
1865) is redescribed and illustrated. Neotypes are designated for Sunius
platynotus Saulcy, 1865 and Dinusa
jebusaea Saulcy, 1865.
Lectotypes are designated for Dinusa hierosolymata Saulcy,
1865, D. particeps Abeille de
Perrin, 1896, and D. angulicollis Fauvel,
1886. The following taxonomic and nomenclatural acts are proposed: Quedius
josue Saulcy, 1865 = Q.
troodites Fagel, 1968,
nov.syn.; Dinusa hierosolymata Saulcy,
1865 = D. saulcyi Abeille de Perrin,
1896, nov.syn.; Dinusa jebusaea Saulcy, 1865 = D. particeps Abeille de Perrin, 1896, nov.syn.; Notothecta hipponensis
(Fauvel, 1886), nov.comb. (ex Dinusa).
Dinusa santschii Wasmann,
1912 from Tunisia is excluded from Dinusa and tentatively moved to Cypha Leach, 1819. Dinusa
now contains eight myrmecophilous species associated with harvester ants
(Messor spp.) and distributed in the East Mediterranean from southern
Greece across Turkey to the Middle East. A checklist of the named species of Dinusa
is provided.
431:
A
revision of Pronomaea Erichson.
II. A new species from China and additional records (Coleoptera: Staphylinidae:
Aleocharinae).
Additional records of four species of Pronomaea Erichson,
1837 are reported from the West Palaearctic region, among them the first record
of P. araxicola Reitter,
1898 from Afghanistan. Pronomaea maxima nov.sp. from Yunnan (China) is
described and illustrated. The genus is now represented in the Palaearctic
region by seventeen species.
432:
A
revision of the Habrocerinae of the world. V. A new species of Habrocerus
from China
and additional records (Coleoptera: Staphylinidae).
Habrocerus
bisetosus nov.sp.
(China: S-Gansu: Qinling Shan) is described, illustrated, and distinguished from
other species of the H. capillaricornis group. Additional records of
seven named and one unnamed species of Habrocerus Erichson,
1839 are reported from the Palaearctic region and Central America (Guatemala),
respectively, among them the remarkable record of a male of H. indicus Assing & Wunderle, 1995 from southeastern Yunnan, China.
Its sexual characters are illustrated. The distributions of H. indicus in
the East Palaearctic region and of Habrocerus in China are mapped. Habrocerus
is now represented by 17 and the Habrocerinae by 23 named species worldwide. An
updated world catalogue of Habrocerinae is provided.
433:
On
the Oedichirus fauna of China (Coleoptera: Staphylinidae: Paederinae).
Three micropterous species of Oedichirus Erichson,
1839 from Yunnan, China, are described and illustrated: O. latexcisus
nov.sp. (environs of Dongchuan), O. schuelkei nov.sp. (Diancang Shan),
and O. abbreviatus nov.sp. (environs of Tengchong). The genus is now
represented in China and Taiwan by six described species, four of them
micropterous. A key to the species of China and Taiwan is provided. The
distributions of the micropterous species are mapped.
434:
A
revision of the Megalopaederus species of Taiwan (Coleoptera:
Staphylinidae: Paederinae).
Megalopaederus
Scheerpeltz, 1957, previously regarded as a distinct genus, is assigned to the
genus Paederus
Fabricius, 1775 as a subgenus. The Megalopaederus
species of Taiwan are revised. In all, eighteen named locally endemic species
belonging to two species groups are recognized, thirteen of them are described
and illustrated for the first time: Paederus
(Megalopaederus)
virgifer
n. sp. (Nantou,
Chiai);
P. (M.)
incurvatus
n. sp. (Kaohsiung);
P. (M.)
adiectus
n. sp. (Pingtung);
P. (M.)
aequilobatus
n. sp. (Pingtung);
P. (M.)
furcispinosus
n. sp. (Nantou);
P. (M.)
excisissimus
n. sp. (Pingtung);
P. (M.)
trihamatus
n. sp. (Taoyuan);
P. (M.)
bivirgatus
n. sp. (Hualien);
P. (M.)
anmamontis
n. sp. (Taichung);
P. (M.)
semiflavus
n. sp. (Taitung);
P. (M.)
alesi
n. sp. (Hualien);
P. (M.)
discurrens
n. sp. (Taichung);
P. (M.)
incisus
n. sp. (Nantou). Rescriptions and illustrations of the five previously described
species, as well as of P.
(M.)
lewisi
Cameron, 1930 from Japan and P.
biacutus
Li et al., 2013 from mainland China
are
provided, and P.
(M.)
bucculentus
n. sp. (Japan: Honshu: Nagano Prefecture) is described. The latter three species
were included in the revision to tentatively assess the phylogenetic
affiliations of Taiwanese Megalopaederus.
Paederus
fulvocaudatus Adachi,
1939, previously considered a subspecies of P.
kosempoensis
Bernhauer, 1922, represents a distinct species. A lectotype is designated for P.
kosempoensis
Bernhauer, 1922. Additional material composed exclusively of females and
probably representing six undescribed species is listed. The available
information on the natural history of Taiwanese Megalopaederus
is compiled. A key to the named species of Taiwan is provided. Their
distributions are mapped.
435:
On
the Staphylinidae (Coleoptera)
of Crete II. Seven new species, a new synonymy, and additional records.
Approximately
110 species of Staphylinidae collected in 2013 and 2014 are reported from the
Greek island Crete. Seven species of Aleocharinae (four species), Paederinae
(one species), and Staphylininae (two species) are described and illustrated,
five of them are probably local endemics: Bellatheta
albimontis
n. sp. (Lefka
Ori); B.
idana
n. sp. (Psiloritis); Geostiba
(Sipalotricha)
paulexcisa
sp n. (Psiloritis); Myrmecopora
(Myrmecopora)
thriptica
n. sp. (Orno
Thriptis; associated with Messor
sp.); Sunius
thripticus
n. sp. (Orno
Thriptis); Quedius
praecisus
n. sp. (Orno
Thriptis, but probably more widespread in Crete); Xantholinus
erinaceus
n. sp. (several localities). The sexual characters of some previously described
species are illustrated. The following synonymy is proposed: Quedius
nemoralis
Baudi di Selve, 1848 = Q.
candicus
Coiffait, 1976, syn. n. The genus Bellatheta
Roubal, 1928 is reported from Crete for the first time. Some previous records
are rectified. The diversity and biogeography of the endemic species of Crete is
discussed. A revised list of the 67 currently known named endemic (sub-)species
of Staphylinidae of Crete is provided.
436:
Three
new species of Domene from China
(Coleoptera: Staphylinidae: Paederinae).
Three
species of Domene Fauvel,
1873 from China, are described and illustrated: Domene (Macromene)
affimbriata sp. n. (southeastern Yunnan: Dawei Shan near Pingbian), D.
(M.) praefigens sp. n. (eastern Yunnan: region to the north,
northwest, and northeast of Kunming), and D. (M.) emeiana
sp. n. (Sichuan: Emei Shan). The distributions of the new species and other
named species of the genus in China and Taiwan are mapped. Domene now
includes 64 named species, 15 of which have been recorded from China (13) and
Taiwan (2).
437:
New
species and additional records of Lathrobium
and Elytrobium from the Palaearctic region, with special reference to the
fauna of East Yunnan (Coleoptera:
Staphylinidae: Paederinae).
Seventeen
species of Lathrobium Gravenhorst,
1802 and one of Elytrobium Assing,
2013 are (re-)described and illustrated: Lathrobium struyvei sp. n.
(Spain), a close relative of L. dimidiatipenne Bernhauer,
1910; L. kiruense sp. n. (Nepal: Annapurna) of the L. emodense
group; L. undosum sp. n. (Nepal: Solu Khumbu), tentatively assigned to
the L. discissum group; L. crenatum sp. n. (China: East Yunnan), L.
rostratum sp. n. (China: East Yunnan), L. tricarinatum sp. n. (China:
East Yunnan), and L. laciniatum sp. n. (China: East Yunnan), all four of
them of the L. tentaculatum group; L. daweianum sp. n. (China:
Southeast Yunnan) of the newly established L. daweianum group; L.
coadultum sp. n. (China: Southeast Yunnan) of the newly established L.
coadultum group; L. grebennikovi sp. n. (China: Shaanxi: Qinling
Shan) and L. cavisulcatum sp. n. (China: Shaanxi: Qinling Shan) of the L.
varisternale group; L. abruptum sp. n. (China: Sichuan: Gongga Shan),
a close relative of L. hailuogouense Peng et al., 2012; L. erectum sp. n. (China: Guizhou:
Fanjing Shan) of the L. fissispinosum group; L. scaphiforme sp. n.
(Japan: Honshu); L. volutum sp. n. (Japan: Honshu); L. trabale sp.
n. (Japan: Honshu); L. oharai Watanabe,
2014; Elytrobium edentulum sp. n. (China: Shaanxi: Qinling Shan). The
diversity, biogeography, natural history, and phylogenetic affiliations of the Lathrobium
fauna of East Yunnan, from where the genus is reported for the first time, are
discussed. The general distribution of Lathrobium and the individual
distributions of the newly described species in Yunnan are mapped. Additional
records of 30 described and one unnamed Lathrobium species are reported
from Middle Asia (one species), Nepal (five species), from the Chinese provinces
Yunnan (five species), Sichuan (five species), Shaanxi (one species), and
Guizhou (one species), from Taiwan (one species), and from Japan (13 species).
Three of these species are reported from Japan for the first time. Additional
records of two Elytrobium species are reported from China and Japan. Lathrobium
is now represented in the Palaearctic region by 563 species and ten subspecies,
in China by 206 species (67 of which are confined to Yunnan), in Japan by 114,
and in the Himalaya by 70 species. Elytrobium currently includes a total
of seven species distributed in China, Taiwan, and Japan.
438:
On
the Staphylinidae of the Greek island Chios (Insecta: Coleoptera).
A study of 536 specimens of Staphylinidae collected in the Greek island Chios in
December 2014 yielded 41 species, among them at least three undescribed
island-endemic species and two first records from Greece. A species list is
provided. Xantholinus (Helicophallus)
chiosicus nov.sp., at present the sole named island-endemic representative
of the family in Chios, is described and illustrated. Geostiba bulbifera Zerche,
1988, nov.syn. is placed in synonymy with G. maxiana (Tikhomirova,
1973). The first discovery of a winged male of Geostiba maxiana confirms
an earlier hypothesis that this species is wing-dimorphic.
439:
A
new species of Phloeocharis Mannerheim
from Spain, with a note on P. brachyptera Sharp (Coleoptera: Staphylinidae: Phloeocharinae).
Phloeocharis
gredosensis
nov.sp. (Central Spain: Sierra de Gredos) is described, illustrated, and
distinguished from other micropterous Phloeocharis species known from
Spain. Material of P. brachyptera Sharp,
1873 from various regions in the Cordillera Cantabria and the western Pyrenees
(northern Spain) is studied. Based on an examination of the male sexual
characters, the previously established synonymy of P. umbratilis Eppelsheim,
1880 with P. brachyptera is confirmed. The male sexual characters of P.
brachyptera and P. microptera Fauvel,
1898 are illustrated. Phloeocharis Mannerheim,
1830 now includes 45 species, twelve of which have been recorded from Spain.
440:
A
revision of Amarochara Thomson
of the Holarctic region V. A new species from China, a new combination, the male
of A. caeca Assing,
and additional records (Coleoptera: Staphylinidae: Aleocharinae: Aleocharini).
Amarochara
daweiana
nov.sp. (China: Yunnan) is described and illustrated. Additional records of
species of Amarochara Thomson, 1858
are reported from the Palaearctic region, among them several first records from
China and Laos. The previously unknown male sexual characters of A. caeca
Assing, 2002 are described and
illustrated. A new combination is established: Ocalea nepalica (Pace,
2013) nov.comb. (ex Amarochara). The genus is now represented in the
Holarctic region by 31 species.
441:
A
revision of Palaearctic and Oriental Rugilus. V. Two new species from
China and additional records (Coleoptera: Staphylinidae: Paederinae).
Two species of Rugilus Leach,
1819 from China are described and illustrated: R. (Rugilus) emeimontis
nov.sp. (Sichuan: Emei Shan), the third endemic Rugilus species from the
Emei Shan, and R. (R.) rugosissimus nov.sp. (Guizhou:
Fanjing Shan), the first record of Rugilus sensu strictu from Guizhou.
Additional records of 15 previously described species are reported from the
Palaearctic and Oriental regions, among them the first record of R. dabaicus
Assing, 2012 from the Qinling Shan
(Shaanxi). Two probably undescribed species from Yunnan and Guizhou, which are
currently represented only by females, remain unnamed. Including the new
species, Rugilus is now represented in the Palaearctic and Oriental
regions by 100 species and one subspecies.
442:
On
Orsunius III. Four new species from China and Thailand, and additional
records (Coleoptera: Staphylinidae: Paederinae: Medonina).
Four
species of Orsunius Assing,
2011 are described and illustrated: O. affimbriatus nov.sp. (China:
Guangdong); O. incurvatus nov.sp. (Thailand); O. granulosissimus
nov.sp. (China: Yunnan); O. confluens nov.sp. (China: Yunnan). Additional
records of four previously described species are reported, among them the first
record of O. cuneatus Assing,
2014 from Thailand. A new key to species and an updated catalogue are provided.
The genus currently comprises 21 named species.
443:
On
the identity of Atheta opacicollis (Fauvel)
(Coleoptera: Staphylinidae: Aleocharinae: Oxypodini).
Types and additional material of Atheta opacicollis (Fauvel, 1878) are revised. A lectotype is designated. The
species is reported from Syria for the first time. Diagnostic characters
distinguishing A. opacicollis from the similar A. meybohmi Assing,
2011 are described and illustrated.
444:
On
the Orphnebius fauna of the East Palaearctic region. VI. Six new species
from China and Taiwan, and additional records (Coleoptera: Staphylinidae:
Aleocharinae: Lomechusini).
Six
species of Orphnebius Motschulsky,
1858 are described and illustrated: Orphnebius dishamatus nov.sp.
(Yunnan), O. incrassatus nov.sp. (Yunnan), and O. formosanus
nov.sp. (Taiwan) of the O. hauseri group; O. tridentatus nov.sp.
(Yunnan) of the O. nanlingensis group; O. (Deroleptus)
multimpressus nov.sp. (Yunnan); O. (Deroleptus)
planicollis nov.sp. (Yunnan). Deroleptus Bernhauer,
1915, previously regarded as a distinct genus, is placed in Orphnebius as
a subgenus. Additional records of nine species from North India, Nepal, and
China are reported. An updated catalogue of the Orphnebius fauna of the
East Palaearctic region and a revised key to the species of China and Taiwan are
provided. Orphnebius is now represented in the East Palaearctic region by
38 named species, 19 of which are distributed in the Himalaya, 18 in China, and
one in Taiwan.
445:
On
the Tetrabothrus fauna of China (Coleoptera: Staphylinidae: Aleocharinae:
Lomechusini).
Five
species of Tetrabothrus Bernhauer,
1915
from China are (re-) described and illustrated: T. cavus nov.sp. (Yunnan,
Shaanxi), T. inflexus nov.sp. (Yunnan), T. brevalatus nov.sp.
(Yunnan), T. rubricollis nov.sp. (North Sichuan), and T. chinensis
Pace, 2012. The six named species
recorded from China thus far are keyed and their distributions are mapped. In
total, the Tetrabothrus material known from China at present amounts to
only twelve specimens, one of them unnamed. Including the new species, the genus
currently includes 28 species, 18 of which have been recorded from the southern
East Palaearctic region.
446:
On
Stilicoderus and Stiliderus IV. Two new species from China and
additional records (Coleoptera: Staphylinidae: Paederinae).
Stilicoderus
hainanus
nov.sp., the first representative of the genus to be recorded from the Chinese
island Hainan, and S. daweianus nov.sp. (China: Yunnan: Dawei Shan) are
described and illustrated. Additional records of eight previously described
species are reported. The distributions of four species in China are mapped. An
updated catalogue of the species of Stilicoderus Sharp, 1889 and Stiliderus Motschulsky, 1858 recorded from the East Palaearctic region,
including Northeast India and Burma, is provided. Stilicoderus now
comprises 107 species, 39 of which have been recorded from the East Palaearctic
region. The country with, by far, the most diverse Stilicoderus fauna is
China (26 species), where as many as 18 species have been recorded from Yunnan
alone.
447:
On
the Harpopaederus fauna of China (Coleoptera: Staphylinidae: Paederinae).
Types
and additional material of Paederus Fabricius,
1775, subgenus Harpopaederus Scheerpeltz,
1957, from China are revised. Seven species are (re-) described and/or
illustrated: Paederus (Harpopaederus)
dangchangensis Li & Zhou,
2007; P. (H.)
lineodenticulatus Li &
Zhou, 2007; P. (H.)
cultellatus nov.sp. (Daba Shan at the border between Chongqing,
Hubei, and Shaanxi); P. (H.)
deplectens nov.sp. (Northwest Sichuan); P. (H.)
edentulus nov.sp. (Gansu: mountains to the southeast of Longnan); P.
(H.) minicus
nov.sp. (Gansu: Min Shan); P. (H.)
chinensis Bernhauer,
1931. Paederus chinensis, the type species of the subgenus Oedopaederus,
is assigned to Harpopaederus, resulting in the synonymy Harpopaederus
Scheerpeltz, 1957 = Oedopaederus
Scheerpeltz, 1957, nov.syn. Paederus
pseudobaudii Aleksandrov,
1934, nov.syn., is placed in synonymy with P. gottschei Kolbe,
1886. The
intraspecific variation of the sexual characters in P. dangchangensis is
illustrated and discussed. Additional records of nine previously described
species are reported. The general distribution of the subgenus in China, as well
as the individual distributions of all the Harpopaederus species recorded
from China are illustrated. A catalogue is provided. Harpopaederus now
includes 20 species, 15 of which have been recorded from China (14 exclusive).
With one exception (P. gottschei),
all the Chinese representatives of the subgenus are distributed in the Qinling
Shan, the Daba Shan, and adjacent mountain ranges, as well as in West Sichuan.
448:
Two
new genera of Medonina from China and Japan (Coleoptera: Staphylinidae:
Paederinae).
Granimedon
nov.gen. (type species: G. anguliceps nov.sp.) and Coimedon nov.gen.
(type species: Lithocharis dissimilis Sharp,
1874) are described and illustrated. The former includes three species, all of
them from southeastern Yunnan, China: Granimedon anguliceps nov.sp.; G.
effeminatus nov.sp.; G. creber nov.sp. Coimedon is represented
only by C. dissimilis (Sharp,
1874), nov.comb., from Kyushu, South Japan. All four species are (re-)described
and illustrated. A key to the species of Granimedon is provided. The
distributions of the Granimedon species are mapped.
449:
On
the Nepalota fauna of China (Coleoptera: Staphylinidae: Aleocharinae:
Athetini).
Types and additonal material of the East Palaearctic aleocharine genus Nepalota
Pace, 1987 are examined. Eleven
species are (re-)described and illustrated: N. franzi Pace,
1987; N. globifera Pace,
1998; N. smetanai Pace,
1998; N. guangdongensis Pace,
2004; N. fellowesi Pace,
2004; N. crocea nov.sp. (Yunnan); N. cuneata nov.sp. (Yunnan); N.
mocytoides nov.sp. (East Yunnan); N. prominula nov.sp. (Yunnan); N.
tuberifera nov.sp. (Yunnan); N. daweiana nov.sp. (Yunnan). Five
synonymies are proposed: N. gansuensis Pace,
1998 = N. qinlingmontis Pace,
2011, nov.syn.; N. globifera Pace,
1998 = N. caluoensis Pace,
2011, nov.syn., = N. daxuensis Pace,
2011, nov.syn.; N. guangdongensis Pace,
2004 = N. rougemonti Pace,
2011, nov.syn. = N. ruficollis Pace,
2011, nov.syn. Nepalota franzi is reported from China for the first time.
In all, 35 valid species are currently known from the East Palaearctic, four of
them of doubtful status (males unknown). Twenty species have been described or
reliably recorded from China exclusive of Taiwan. The diversity is greatest in
Yunnan (fourteen species, seven of them exclusive), followed by Hubei (five
species; three exclusive); Sichuan (four species, one exclusive), Gansu and
Shaanxi (three species each), Guangdong (two species), Zhejiang, Chongqing,
Jiangxi, Guangxi, Guizhou, and Beijing (one species each). The distributions of
15 species are mapped. An updated checklist of Nepalota is provided.
450:
Two
new species of Chinecallicerus (Coleoptera: Staphylinidae: Aleocharinae:
Geostibini).
Chinecallicerus
subater
nov.sp. (China: Northeast Yunnan) and C. reticulatus nov.sp. (China:
Southeast Yunnan) are described, illustrated, and distinguished from their
congeners. An updated key to the species of Chinecallicerus Assing,
2004, a genus currently known only from Yunnan, China, and including six species
is provided. The distributions of the genus and the individual species are
mapped.
451:
A
revision of Porocallus Sharp.
IV. A new species from China and additional records (Coleoptera: Staphylinidae:
Aleocharinae).
Porocallus
cicatricatus
nov.sp. (China: southeastern Yunnan) is
described and illustrated. Additional records of two previously described
species are reported. The currently known distributions of all the species of Porocallus
Sharp, 1888 are mapped. The East
Palaearctic genus now includes seven species.
452:
A
revision of Nazeris VII. Three new species and additional records from
Nepal and China
(Coleoptera: Staphylinidae: Paederinae).
Three
species of Nazeris Fauvel, 1873 are described and illustrated: N.
brevior sp. n. (Nepal: Solu Khumbu) of the N. alticola group; N.
hamulatus sp. n. (China: Shaanxi: Qinling Shan) of the N. longilobatus
group; N. eminens sp. n. (China: Sichuan: Emei Shan), tentatively
assigned to the N. zhangi lineage of the N. cangicus group.
Additional records of eleven described and one unnamed species are reported from
Nepal (four species) and China (eight species). The Palaearctic genus is now
represented by 236 described species (plus seven subspecies), 120 of which are
distributed in mainland China and 49 in the Himalaya.
453:
On
the Staphylinidae of the Greek island Samos (Coleoptera: Staphylinidae).
A
recent field trip to the Greek island Samos yielded 1322 specimens belonging to
at least 117 species of Staphylinidae. Three species are described and
illustrated: Geostiba (Tropogastrosipalia) plicipennis sp.
n. (Oros Ambelos) and Oxypoda (Bessopora) kerkisica sp. n.
(Oros Kerkis) of the Aleocharinae, and Sunius ambelosicus sp. n. (Oros
Ambelos) of the Paederinae. Geostiba plicipennis is the first
representative of the speciose subgenus Tropogastrosipalia Scheerpeltz,
1951 to be recorded from an island. Including the Pselaphinae and Scydmaeninae
recorded by Volker Brachat and Heinrich Meybohm in 2003, as well as previous
literature records, at least 136 species of Staphylinidae are currently known
from Samos, 103 of them named and positively identified, 15 undescribed (five of
the Pselaphinae and ten of the Scydmaeninae), eight tentatively identified, and
at least ten unidentified. Six species are reported from Greece for the first
time. The remarkably diverse staphylinid fauna of Samos includes as many as 18
species that have been recorded exclusively from this island and are most likely
island-endemic, some of them even locally endemic. However, only seven of these
species have been named; the remainder is pending description. Xantholinus
chiosicus Assing,
2015, a species previously hypothesized to be endemic to Chios, is recorded also
from Samos. The diversity of the staphylinid fauna of Samos is evidently greater
than that of other North Aegean islands; it appears to be even greater than that
of the much larger island Rhodes, an observation that may be explained with the
geography, geology, and ecological characteristics of Samos. A comprehensive
list of the named and unnamed species currently known from Samos is provided.
454:
New
species and additional records of Lomechusini from the Palaearctic region,
primarily from China (Coleoptera: Staphylinidae: Aleocharinae).
Six
species of Lomechusini from China and North India are (re-)described and
illustrated: Drusilla yunnanensis Pace,
1993; D. watanabei Maruyama &
Kishimoto, 2002; D. flagellata sp. n. (China: Yunnan); D.
palata sp. n. (China: Yunnan); Amaurodera angularis sp. n. (North
India: West Bengal); Lomechusoides penicillatus sp. n. (China: Gansu).
Two synonymies are proposed: Drusilla canaliculata (Fabricius,
1787) = D. puncticollis Motschulsky,
1845, syn. n.; Amaurodera smetanai Pace,
1992 = A. yaoana Pace, 1992,
syn. n. Additional records of 29 previously described species of Drusilla
Leach, 1819, Pella Stephens,
1835, Amaurodera Fauvel,
1905, Tetrabothrus Bernhauer, 1915,
and Lomechusoides Tottenham,
1939 are reported from the Palaearctic region, primarily from China, among them
several new country records and the first record of Pella lugens (Gravenhorst, 1802) from the East Palaearctic. The previously
unknown aedeagus of Pella reuteri Assing,
2008, the previously unknown female sexual characters of Amaurodera ilamica
Assing, 2005, and the primary
sexual characters of several Pella species recorded from China are
illustrated. The distributions of the Drusilla species currently known
from China are illustrated.
455:
A
revision of Neosclerus V. Thirteen new
species and additional records (Coleoptera: Staphylinidae: Paederinae).
Thirteen
species of Neosclerus Cameron,
1924 are described and illustrated: N. albens sp. n. (Sri Lanka), N.
areolatus sp. n. (Thailand), N. biangulatus sp. n. (Thailand), N.
daweianus sp. n. (China: Yunnan), N. discolor sp. n. (Burma,
Thailand), N. figens sp. n. (China: Yunnan), N. forficatus sp. n.
(Sri Lanka), N. fractus sp. n. (India: Meghalaya), N. khasicus sp.
n. (India: Meghalaya), N. revolutus sp. n. (South India), N.
sagittatus sp. n. (India: Meghalaya), N. semicalvus sp. n.
(Thailand), and N. trisinuatus sp. n. (China: Yunnan). The previously
unknown male sexual characters of N. granulicollis Cameron,
1924 are described and illustrated. Additional records of twelve named and three
unnamed species are reported, among them several new country records. The genus
now comprises 44 named species in five species groups, one of them newly
established. The countries with the greatest diversity are China, India, and
Thailand (ten species each), and Taiwan (nine locally endemic species). The
distribution of the genus as a whole, as well as the individual distributions of
34 species are mapped. An updated catalogue is provided.
456:
A
revision of Sunius XV. Six new species from Kyrgyzstan and the Himalaya,
and additional records (Coleoptera: Staphylinidae: Paederinae).
Six
micropterous species of Sunius Stephens,
1829 are described and illustrated: S. alaianus sp. n. (Kyrgyzstan) of
the S. viator group; S. marratus sp. n. (North India: West
Bengal), S. virilis (North India: West Bengal), S. deplectens sp.
n. (North India: West Bengal), S. dearmatus sp. n. (East Nepal), and S.
migrus sp. n. (Central Nepal), all five of them of the S. manasluensis
group. Additional records of 16 species are reported both from the West and the
East Palaearctic regions. The genus is now represented in the Palaearctic region
by 126 described species and two subspecies.
457:
The
first species of Borboropora KRAATZ
from
the East Palaearctic region (Coleoptera: Staphylinidae: Aleocharinae: Falagriini).
Borboropora
indica
nov.sp. (India: Arunachal Pradesh), the first representative of the genus
recorded from the East Palaearctic region, is described and illustrated. Borboropora
pseudoquadriceps Delgado &
Santiago-Jimιnez, 2009 and B. mixe Delgado & Santiago-Jimιnez, 2009 from Mexico are
excluded from Borboropora Kraatz,
1862 and tentatively moved to Anaulacaspis Ganglbauer, 1895 and Falagrioma Casey,
1906, respectively, resulting in the binomina Anaulacaspis pseudoquadriceps
(Delgado & Santiago-Jimιnez, 2009), nov.comb., and Falagrioma
mixe (Delgado & Santiago-Jimιnez,
2009), nov.comb. Thus, Borboropora currrently includes a total of seven
species..
458:
A
new myrmecophilous species of Eurysunius from Turkey (Coleoptera:
Staphylinidae: Paederinae).
Astenus
(Eurysunius)
kociani
nov.sp. (Turkey: Nevşehir) is described, illustrated, and distinguished
from other species of Eurysunius Reitter,
1909 known from Turkey, particularly the similar and evidently closely allied Astenus
sexsetosus Assing, 2002. The
species was found associated with an unidentified species of Tetramorium Mayr,
1855.
459:
New
species and additional records of Dolicaonina from Ecuador (Coleoptera:
Staphylinidae: Paederinae).
Five species of Gnathymenus Solier,
1849 are described and illustrated, all of them from Ecuador: Gnathymenus
penicillatus nov.sp. (Cotopaxi); G. crucifer nov.sp. (Zamora
Chinchipe); G. stipulatus nov.sp. (Zamora Chinchipe); G. abscisus
nov.sp. (Zamora Chinchipe); G. multiplicus nov.sp. (Esmeraldas). One of
these species (G. penicillatus) is micropterous, the remainder
macropterous. Additional records of three species of Gnathymenus and one
of Stenopholea Herman,
1969 are reported from Ecuador.
460:
A
revision of the Lithocharis species of the Palaearctic, Oriental and
Australian regions (Coleoptera: Staphylinidae: Paederinae: Medonina).
The
species of the medonine genus Lithocharis Dejean, 1833 of the Palaearctic (Mauritania included),
Oriental, and Australian regions are revised. In all, 23 species are recognized,
one of them of doubtful identity (male and type depository unknown) and one
(male unknown) of doubtful generic assignment. Except for few, partly previously
revised species from the West Palaearctic region, all the species are (re-)described
and illustrated, six of them for the first time: Lithocharis socotrana
nov.sp. (Yemen: Socotra); L. bitriangulata nov.sp. (Sumatra); L.
inermis nov.sp. (Thailand); L. latexcisa nov.sp. (Nepal); L.
ornatrix nov.sp. (Malaysia, Indonesia); L. parvincisa nov.sp. (Indonesia:
Sulawesi). Lithocharis erythroptera Gemminger
& Harold, 1868 is revalidated. Five synonymies are established: L.
erythroptera Gemminger & Harold,
1868 = L. penicillata Cameron,
1928, nov.syn.; L. nigriceps Kraatz,
1859 = L. changlingensis Li,
1992, nov.syn.; L. vilis Kraatz,
1859 = L. jacobsoni Cameron,
1928, nov.syn., = L. vexans Cameron,
1936, nov.syn.; Panscopaeus lithocharoides (Sharp, 1874) = Medon subopacus Bernhauer,
1907, nov.syn. Lithocharis gigantea Fauvel,
1878 is excluded from Lithocharis. Lithocharis ochracea (Gravenhorst,
1802) is confirmed as a valid name and removed from the previously proposed
synonymy with Staphylinus tricolor Fabricius,
1787, whose valid present combination is Xantholinus tricolor (Fabricius,
1787). Lectotypes are designated for Lithocharis sororcula Kraatz,
1859, L. fuscipennis Kraatz,
1859, L. penicillata Cameron,
1928, L. uvida Kraatz, 1859,
L. lamellifera Cameron,
1928, L. distinguenda Cameron,
1928, L. sordida Cameron,
1928, L. timorensis Cameron,
1928, Medon subopacus Bernhauer,
1907, and Staphylinus tricolor Fabricius,
1787. Inhabitants of decaying organic matter and active flyers, Lithocharis
species are generally remarkably widespread. Males of this genus are
occasionally subject to conspicuous teratologies (aedeagus completely reduced;
male secondary characters absent). A key to the Lithocharis species of
the Palaearctic (including Mauritania), Oriental, and Australian regions, and a
catalogue are provided.
461:
A
revision of Trisunius V. Three new species and additional records,
including the first records of the genus from Vietnam (Coleoptera: Staphylinidae:
Paederinae).
Three species of the medonine genus Trisunius
Assing, 2011 from North Thailand
are described and illustrated: Trisunius
scaphiformis nov.sp. (Doi Inthanon), T. conlectus nov.sp. (Doi
Inthanon), and T. penicillatus nov.sp.
(Doi Pha Hom). Additional records of four species are reported from East Yunnan,
Thailand, and Vietnam. The discovery of two species in Vietnam represents the
first record of the Trisunius from this country. The distribution of the
genus as a whole and the individual distributions of three widespread species
are mapped. Trisunius now includes 22 species.
462:
First
record of Pseudomedon from the Oriental Region (Coleoptera: Staphylinidae:
Paederinae: Medonina).
Pseudomedon
discolor nov.sp.
(North Vietnam: environs of Sa Pa), representing the first record of the genus
from the Oriental region and from Vietnam, is described and illustrated. An
additional record of Pseudomedon
kazakhstanicus Assing, 2008 is
reported from Kazakhstan. The genus now includes a total of 23 species.
463:
A
revision of Sclerochiton. II. Two new species and additional records,
including the first records of the genus from Vietnam (Coleoptera: Staphylinidae:
Paederinae).
Two species of Sclerochiton Kraatz,
1859 are described and illustrated: Sclerochiton
pectinatus nov.sp. (Laos) and S.
peniculatus nov.sp. (North Vietnam). Additional records of ten species are
reported. The genus is recorded from Vietnam for the first time (two species).
The distribution of the genus in the southern East Palaearctic and Oriental
regions and the individual distributions of all the species treated are mapped. Sclerochiton
now includes a total of 23 described species, 17 of which are distributed in the
southern East Palaearctic and Oriental regions.
464:
A
revision of the Habrocerinae of the world. VI. The first records of the
subfamily from Vietnam (Coleoptera: Staphylinidae).
Habrocerus
wunderlei nov.sp.
(North Vietnam: environs of Sa Pa) is described, illustrated, and distinguished
from other species of the H. schwarzi group. Additional records of two
species are reported from North Vietnam and Thailand. The subfamily Habrocerinae
is reported from Vietnam for the first time. Habrocerus Erichson,
1839 is now represented by 18 and the Habrocerinae by 24 named species worldwide.
A revised and updated world catalogue of subfamily is provided.
465:
Three
new species of Deinopteroloma from Vietnam and China (Coleoptera:
Staphylinidae: Omaliinae).
Three species of Deinopteroloma Jansson,
1946 are described and illustrated: Deinopteroloma
uncinatum nov.sp. (North Vietnam: environs of Sa Pa), D. dispar nov.sp. (North Vietnam: environs of Sa Pa), and D.
obtortum nov.sp. (China: southern Gansu). The genus is reported from Vietnam
and the Oriental region for the first time. Additional records of D. chiangi
Smetana, 1990 and D. gracile Smetana, 2001 are reported from Taiwan and Sichuan,
respectively. The distribution of the genus in Asia east of the Himalaya is
mapped. Including the new species, Deinopteroloma currently includes 17
species and is distributed in the western Nearctic, the southern East
Palaearctic, and the northern Oriental regions.
466:
A
revision of Porocallus SHARP.
V. The first records of the genus from Vietnam (Coleoptera: Staphylinidae:
Aleocharinae).
Porocallus
vietnamensis
nov.sp. (North Vietnam: environs of Sa Pa) is
described and illustrated. Porocallus wrasei Assing,
2012, a species previously known only from the Chinese province Yunnan, is
reported from Vietnam for the first time. The East Palaearctic genus Porocallus
Sharp, 1888 now includes eight
species.
467:
A
revision of Othiini. XIX. Two new species of Othius, the first record of
the genus from Vietnam, and additional records (Coleoptera: Staphylinidae:
Staphylininae).
Othius
bihamatus
nov.sp. (Pakistan: North-West Frontier Province), a close relative of O.
sinuosus Assing, 1998, and O.
auster nov.sp. (North Vietnam), the first representative of the genus from
the Oriental region and the southernmost record of the genus known at present,
are described and illustrated. Additional records of 21 species of Othius
Stephens, 1829 and two of Atrecus
Jacquelin du Val, 1856 are reported,
among them the first record of Atrecus brevicornis Smetana,
1967 since the original description. The distribution of Othius in the
East Palaearctic region east of the Himalaya and the individual distributions of
eight species are mapped. Othius and the Othiini now include a total of
128 and 145 species and subspecies, respectively.
468:
A
new species of Algon from Vietnam (Coleoptera: Staphylinidae:
Staphylininae).
Algon
fansipanicus nov.sp.
(North Vietnam: environs of Sa Pa), a micropterous species of the A.
kaiserianus group, is described and illustrated. The A. kaiserianus
group is recorded from Vietnam for the first time.
469:
On
the Lomechusini fauna of Vietnam (Coleoptera: Staphylinidae: Aleocharinae).
Five
species of Lomechusini from the environs of Sa Pa, North Vietnam, are described
and illustrated: Amaurodera projecta nov.sp., Drusilla perforans
nov.sp., Orphnebius fansipanicus nov.sp., Peltodonia atripalpis
nov.sp., and Zyras (Zyras) fansipanicus nov.sp. Three
probably undescribed species remain unnamed for want of males. Type material of
nine described species is revised and illustrated. The previously unknown male
sexual characters of Peltodonia chinensis (Pace,
1998) and Tetrabothrus bicolor Cameron,
1939 are figured for the first time. Two synonymies and two new combinations are
proposed: Tetrabothrus japonicus
Nakane,
1991 = T. vietnamiculus Pace,
2013; nov.syn.; Zyras neoparageminus Hlavαč,
Newton
& Maruyama,
2011 = Z. subgeminus Pace,
2012, nov.syn.; Zyras
(Zyras)
funestus
(Dvořak,
1996), nov.comb. (ex Orphnebius Motschulsky,
1858); Zyras (Glossacantha)
laocaiensis
Pace,
2008, nov.comb. (ex Pella
Stephens,
1833).
A lectotype is designated for Tetrabothrus bicolor Cameron,
1939. Three species and three genera are recorded from Vietnam for the first
time. The Lomechusini fauna of Vietnam is currently represented by 29 species,
partly of doubtful status, in seven genera. A checklist of the Lomechusini of
Vietnam is provided.
470:
A
revision of Zyras Stephens
sensu strictu of China, Taiwan, and Hong Kong, with records and
(re-)descriptions of some species from other regions (Coleoptera: Staphylinidae:
Aleocharinae: Lomechusini).
The
species of the nominal subgenus of the lomechusine genus Zyras
Stephens, 1835 of China, Taiwan, and Hong Kong are revised. In all, 50 named
species, two of them of doubtful identity, are recognized, (re-)described, and
illustrated, among them 33 new taxa: Zyras
(Zyras) athetoides
n. sp. (Sichuan), Z.
(Z.) atrapicalis
n. sp. (Yunnan),
Z.
(Z.) atronitens
n. sp. (Tibet), Z.
(Z.) bangmaicus
n. sp. (Yunnan), Z.
(Z.) bicoloricollis
n. sp. (Yunnan), Z.
(Z.) bisinuatus
n. sp. (Yunnan), Z.
(Z.) caloderoides
n. sp. (Yunnan), Z.
(Z.) dabanicus
n. sp. (Qinghai), Z.
(Z.) discolor
n. sp. (Fujian), Z.
(Z.) exspoliatus
n. sp. (Guangxi), Z.
(Z.) extensus
n. sp. (Yunnan), Z.
(Z.) firmicornis
n. sp. (Fujian), Z.
(Z.) flexus
n. sp. (Fujian), Z.
(Z.) formosanus
n. sp. (Taiwan), Z.
(Z.) gilvipalpis
n. sp. (Yunnan), Z.
(Z.) granapicalis
n. sp. (Sichuan), Z.
(Z.) hebes
n. sp. (Taiwan), Z.
(Z.) inexcisus
n. sp. (China: Gansu, Qinghai;
Russia: Far East, East Siberia), Z.
(Z.) lativentris
n. sp. (Yunnan),
Z.
(Z.) maculicollis
n. sp. (Yunnan), Z.
(Z.) nigrapicalis
n. sp. (Yunnan, Sichuan,
Jiangxi, Taiwan), Z.
(Z.) nigricornis
n. sp. (Hubei, Gansu, Shaanxi,
Sichuan, Qinghai), Z.
(Z.) nigronitens
n. sp. (Yunnan), Z.
(Z.) pulcher
n. sp. (Gansu, Sichuan), Z.
(Z.) rectus
n. sp. (Yunnan), Z.
(Z.) rufapicalis
n. sp. (Taiwan), Z.
(Z.) rufoterminalis
n. sp. (Hubei,
Sichuan),
Z.
(Z.) schuelkei
n. sp. (Fujian, Sichuan),
Z.
(Z.) subobsoletus
n. sp. (Sichuan), Z.
(Z.) tenebricosus
n. sp. (Sichuan, Tibet), Z.
(Z.) tenuicornis
n. sp. (Taiwan), Z.
(Z.) tumidicornis
n. sp. (Taiwan),
Z.
(Z.) volans
n. sp. (Taiwan). Previous
records of five species from China are considered doubtful. As many as 19
species remain unnamed for want of mature males. A catalogue and a key to
species are provided. The distributions of 46 species are mapped. Additional
records and (re-)descriptions of 15 species from other Palaearctic and
Oriental regions are provided. Two species are described for the first time:
Z.
(Z.) iniquus
n. sp. (Pakistan) and Z.
(Z.) articollis
n. sp. (Laos). Seven synonymies
and a new combination are proposed: Zyras
beijingensis
Pace, 1993 = Z. restitutus
Pace, 1993, n. syn.; Zyras
alboantennatus
Pace, 1986 = Z. sichuanorum
Pace, 2012, n. syn.; Z.
birmanus
Scheerpeltz, 1965 = Z.
pseudobirmanus
Scheerpeltz, 1965, n. syn.; Z.
hongkongensis
Pace, 1999 = Z. benenensis
Pace, 2001, n. syn.; Z.
kambaitiensis
Scheerpeltz, 1965 = Z.
ferrugineiventris
Scheerpeltz, 1965, n. syn., = semiasperatus
Scheerpeltz, 1965, n. syn.; Z.
wei
Pace, 1993 = Z. qingchengensis
Pace, 2012, n. syn.; Zyras
brignolii
(Pace, 1986), n. comb. (ex Drusilla).
Lectotypes are designated for Zyras
chinkiangensis
Bernhauer, 1939 and Z.
seminigerrimus
Bernhauer, 1933.
471:
On
some Lomechusini of the Palaearctic and Oriental regions (Coleoptera:
Staphylinidae: Aleocharinae).
Types
and additional material of twelve genera, one of them new, of Lomechusini from
the Palaearctic and Oriental regions are revised. In all, 70 species are
(re-)described and/or illustrated, 51 of them new: Orphnebius bakeri Bernhauer, 1929; O. breviceps Cameron, 1946; O. opticus Cameron, 1946; O. bicuspis sp. n. (Laos; India:
Arunachal Pradesh); O. biformis sp. n. (Indonesia: Sumatra); O.
cernens sp. n. (Laos); O. dilatatus sp. n. (Laos); O. effeminatus
sp. n. (Malaysia); O. extensus sp. n. (Laos); O. fodens sp. n.
(Malaysia: Sabah); O. fuscapicalis sp. n. (Laos); O. fusicollis
sp. n. (Laos); O. grandicollis sp. n. (Laos); O. integer sp. n.
(Laos); O. latitibialis sp. n. (Laos); O. lunatus sp. n. (Laos); O.
nigrapicalis sp. n. (Laos); O. reductus sp. n. (Laos); O. retunsus
sp. n. (Laos); O. serratus sp. n. (Laos); O. spinans sp. n.
(India: Arunachal Pradesh); Orphnebius (Deroleptus) falagrioides
Bernhauer, 1929; O. (D.)
laticeps Cameron, 1925;
O. (D.) multimpressus Assing,
2015; O. (D.) niger (Cameron,
1939); O. (D.) siamensis Cameron,
1939; O. (D.) baccillatus sp. n. (Laos); O. (D.)
biimpressus sp. n. (Indonesia: Sumatra); O. (D.) carinatus
sp. n. (Laos); O. (D.) cultellatus sp. n. (Thailand, Laos);
O. (D.) discrepans sp. n. (China: Yunnan); O. (D.)
dispar sp. n. (India: Arunachal Pradesh); O. (D.) gracilior
sp. n. (India: Arunachal Pradesh); O. (D.) septemcuspis sp.
n. (Laos); O. (D.) sexcarinatus sp. n. (Indonesia:
Sumatra); O. (D.) spoliatus sp. n. (Laos); O. (D.)
tortus sp. n. (India: Meghalaya); O. (D.) triapicalis
sp. n. (China: Sichuan); O. (D.) ulcerosus sp. n.
(Malaysia: Sabah); O. (D.) vates sp. n. (Laos); Drusilla
bifida sp. n. (Thailand); Rabdotodrusilla pectinata sp. n.
(Thailand); Amaurodera meorum Pace,
1992; A. thailandensis Pace,
1986; A. arunica sp. n. (Nepal); A. dentata sp. n. (Thailand); A.
disparicollis sp. n. (Indonesia: Sumatra); A. fasciata sp. n.
(Thailand); A. gilvios sp. n. (Thailand); A. parvoculata sp. n.
(Thailand); A. reticulata sp. n. (Thailand); A. spinans sp. n.
(Indonesia: Sumatra); A. varicollis sp. n. (Indonesia: Sumatra); Tetrabothrus
borneensis Cameron, 1943; T.
indicus Cameron,
1939; T.
inflexus Assing,
2015; T.
neoguineensis
Pace, 2012; T. pubescens Bernhauer,
1915; T. breviatus sp. n. (China: Sichuan); T. collucatus sp. n.
(Laos); T. nilgiricus sp. n. (India: Tamil Nadu); T. punctiventris
sp. n. (Malaysia: Sabah); T. sulawesicus sp. n. (Indonesia: Sulawesi
Utara); Zyras (Zyras) gibbus Pace,
2010; Z. (Z.) illecebrosus Last,
1982; Z. (Z.) quasar
Dvořak,
1996; Z. (Z.) porrectus sp. n. (China: Sichuan); Z. (Z.)
wunderlei sp. n. (Indonesia: Bali); Pedinopleurus notabilis (Silvestri,
1946); Aenictoides gen. n. derivata sp. n. (Thailand). The
intrageneric affiliations of Orphnebius Motschulsky,
1858 and the subgeneric concept currently in use are discussed, and new species
(sub-)groups are proposed. The following synonymies are proposed: Deroleptus
Bernhauer, 1915 = Megalocephalobius
Bernhauer, 1929, syn. n.; Orphnebius
breviceps Cameron, 1946 = O.
vorax Pace, 2000, syn. n.; Orphnebius
niger (Cameron, 1939) = O.
turensis Pace, 2012, syn. n.; Drusilla
canaliculata (Fabricius, 1787) = Myrmedonia
polyporina
Gistel,
1857,
syn. n.; Drusilla erichsoni (Peyron,
1857) = Myrmedonia aptera
Peyron,
1858,
syn. n.; Drusilla zyrasoides
Dvořak,
1988 = D. truncatella
Pace,
2004,
syn. n.; Amaurodera kraepelini Fauvel,
1905 = A. rougemonti Pace, 1987,
syn. n., = A. smetanai Pace, 1992,
syn. n., = A. yaoana Pace, 1992,
syn. n.; Tetrabothrus
bicolor Cameron,
1939 = T.
chinensis Pace,
2012,
syn. n.; Tetrabothrus
puetzi Assing,
2009 = T.
taiwanensis Pace,
2010,
syn. n.; Tetrabothrus
clavatus Bernhauer,
1915 = T.
quadricollis Cameron,
1950,
syn. n., = T.
japonicus Nakane,
1991,
syn. n., = T.
vietnamiculus Pace,
2013,
syn. n., = T.
rubricollis Assing,
2015,
syn. n.; Zyras
fratrumkadooriorum
Pace, 1998 = Z.
chumphonensis
Pace,
2004, syn. n.; Pedinopleurus notabilis (Silvestri, 1946) = P. setosicauda Dvořαk,
1996, syn. n. Lectotypes are designated for Deroleptus niger Cameron,
1939, Orphnebius falagrioides Bernhauer,
1929, Myrmedonia aptera
Peyron,
1858, and
Tetrabothrus
indicus Cameron,
1939. Orphnebius
krypticola
Pace, 2007 is designated as the
correct original spelling. Updated catalogues of Amaurodera Fauvel,
1905 and Tetrabothrus Bernhauer,
1915, as well as keys to the Orphnebius species of Laos and to the
Amaurodera species of Thailand are provided. Additional records of 41
named and several unnamed species are reported, among them numerous new country
and province records. The distributions of Amaurodera kraepelini and Tetrabothrus
clavatus are mapped.
472:
Two
new species and additional records of Domene from China and Vietnam
(Coleoptera: Staphylinidae: Paederinae).
Two
species of Domene Fauvel,
1873 from China are described and illustrated: Domene (Macromene) contiger
sp. n. (Guangxi) and D. (M.) exicta sp. n. (Hubei).
Additional records of three species are reported from China (Hebei, Hubei,
Shaanxi, Sichuan) and North Vietnam. Including the new species, Domene
now includes 67 named species, 16 of which have been recorded from China, two
from Taiwan, and two from Vietnam.
473:
Six
new species, a new name, and additional records of Lathrobium from the
Palaearctic region (Coleoptera: Staphylinidae: Paederinae).
Six species of Lathrobium Gravenhorst,
1802 are described and illustrated: Lathrobium kociani nov.sp.
(Kazakhstan), L. retunsum nov.sp. (East Nepal) of the L. nepalense
group, L. unguiferum nov.sp. (Nepal: Dhaulagiri) of the L. emodense
group, L. spinans nov.sp. (Nepal: Manaslu) of the L. pectinatum
group, L. jiajinum nov.sp. (China: Sichuan) of the L. bibaculatum
group, and L. wolongicum nov.sp. (China: Sichuan) of the L.
bibaculatum group. The previously unknown female sexual characters of L.
makaluicum Assing, 2013 are
illustrated. The preoccupied name Lathrobium abruptum Assing,
2015 is replaced with the nomen novum L. novabruptum. Additional records
of 21 species are reported, particularly from Nepal and China. The genus is now
represented in the Palaearctic region by 579 species and nine subspecies. The
Himalayan fauna currently includes 73 species, all of them micropterous and
locally endemic. As many as 211 named species have been recorded from China.
474:
On
Oxypoda besucheti Focarile
(Coleoptera: Staphylinidae: Aleocharinae: Oxypodini).
Based on a study of type material and numerous additional specimens from
several localities in southern Switzerland (Ticino, Grisons) and northern Italy
(Lombardia: Alpi Orobie) a new combination and a new synonymy are proposed Tectusa
besucheti (Focarile, 1982),
nov.comb. (ex Oxypoda Mannerheim,
1830) = T. orobiana Assing,
2012, nov.syn. are proposed. The intraspecific variation of the aedeagal
morphology is described, discussed, and illustrated. The distribution of T.
besucheti is mapped. The habitat of this alpine species is described and
illustrated.
475:
A
revision of Geostiba of the West Palaearctic region. XXII. Two new species from
Jordan and the Caucaus, and additional records (Coleoptera: Staphylinidae:
Aleocharinae).
Two species of Geostiba Thomson,
1858 from the West Caucasus and Jordan are described and illustrated: Geostiba
(Sibiota) articarinata nov.sp. (Russia: Krasnodarskiy Kray) and G.
(Sipalotricha) jordanica nov.sp., the first representative of the
genus to be recorded from Jordan. Additional records of 14 species from the West
Palaearctic region are reported.
476:
A
new species of Tetrasticta from Laos (Coleoptera: Staphylinidae:
Aleocharinae).
Tetrasticta
laotica
nov.sp. from Laos is described, illustrated, and distinguished from its
congeners.
477:
On
the Staphylinidae of the Greek island Karpathos (Insecta: Coleoptera).
A study of 931 specimens of Staphylinidae collected in the Greek island
Karpathos in December 2015 and January 2016 yielded 59 species, among them eight
undescribed island-endemic species and three first records from Greece. A
checklist including also previously recorded species is provided. The currently
known fauna of Karpathos is composed of 69 species of Staphylinidae and shows
stronger affinities to the fauna of Turkey and the Aegean islands off the
Turkish coast than to Crete and mainland Greece (including the Pelopσnnisos).
Ten species (14.5 %) are island endemics, three of which (two of the Pselaphinae
and one of the Scydmaeninae) are undescribed. Two myrmecophilous species that
were previously considered island endemics of Crete and Rhodos, respectively,
are present also in Karpathos. Oxypoda (Atlantoxypoda) bicornuta Assing
nov.sp. of the Aleocharinae, Medon carpathicus Assing
nov.sp. of the M. petrochilosi subgroup (Paederinae), as well as Cephennium
(Phennecium) kerpense Meybohm
nov.sp., Euconnus (Tetramelus) kerpensis Meybohm
nov.sp., and Stenichnus (Stenichnus) kerpensis Meybohm
nov.sp. of the Scydmaeninae are described and illustrated. In an appendix, Oxypoda
(Atlantoxypoda) bistirpata Assing
nov.sp. (southern Anatolia), a close relative of O. bicornuta, is
described and illustrated.
478:
On
the Carabidae of the Greek island Karpathos (Insecta: Coleoptera).
A study of 176 specimens of Carabidae collected in the Greek island Karpathos
in December 2015 and January 2016 yielded 17 species, among them a new
island-endemic species and three first records from Karpathos. The currently
known fauna of Karpathos is composed of 48, that of the Karpathos archipelago of
52 species of Carabidae. Three species are island endemics; the status of one
taxon is uncertain.
479:
On
the Staphylinidae of Turkey XI. Two new species, new synonymies, and additional
records (Insecta: Coleoptera).
Two species of Staphylinidae from Turkey are described and illustrated: Liogluta
nigrobusta nov.sp. (Artvin) of the Aleocharinae and Vulda substricta
nov.sp. (Sinop) of the Xantholinini (Staphylininae). Two synonymies are
proposed: Xenochara Mulsant &
Rey, 1874 = Rheochara Mulsant
& Rey, 1875, nov.syn.; Xantholinus luteipennis Coiffait,
1970 = X. khachikovi Anlaş,
2014, nov.syn. Additional records of 19 named and one unnamed species are
reported, several of them considerably expanding the previously known ranges and
seven of them representing first records from Turkey.
480:
On
Stilicoderus and Stiliderus V. Two new species, additional
records, and a catalogue (Coleoptera: Staphylinidae: Paederinae).
Stilicoderus
confusus
nov.sp. (China: Yunnan; India: Meghalaya) and S. brachypterus nov.sp.
(East Nepal), two species that had previously been confounded with S.
granulifrons (Rougemont, 1985),
are described and illustrated. The distributions of the species of the S.
granulifrons group are mapped. Additional records of eleven species of Stilicoderus
Sharp, 1889 and four of Stiliderus
Motschulsky, 1858 are reported,
among them new records from Thailand and Vietnam. A comprehensive catalogue of
the 111 species of Stilicoderus and 52 species of Stiliderus
currently known is provided.
481:
A
revision of Nazeris VIII. Five new species from China and additional
records (Coleoptera: Staphylinidae: Paederinae).
Five species of Nazeris Fauvel,
1875 from China are described and illustrated: Nazeris bihamatus nov.sp.
(SW-Guangxi), N. latilobatus nov.sp. (N-Guangxi: Maoer Shan), N.
obtortus nov.sp. (N-Guangxi: Huaping), N. clavator nov.sp.
(Guangdong), and N. latibasalis nov.sp. (Sichuan). Nazeris clavator
represents the first record of the genus from Guangdong province. Additional
records of 23 described and one undescribed species are reported from the
Caucasus, the Himalaya, Vietnam, and China. The genus now includes a total of
241 described species and seven subspecies. It is represented in China by 125
described species.
482:
On
some Acanthoglossa and Hypomedon species II. Two new species, a
new synonymy, and additional records (Coleoptera: Staphylinidae: Paederinae).
Two species of Hypomedon Mulsant
& Rey, 1878 are described and illustrated: H. nasutus nov.sp.
(Thailand) and H. bicornutus nov.sp. (Laos). Based on a study of types,
the following synonymy is proposed: Acanthoglossa
hirta Kraatz; 1859 = Medon
sutteri Scheerpeltz, 1957 nov.syn. Records of Hypomedon
debilicornis (Wollaston, 1857)
and H. galilaeus (Bordoni,
1980) are reported, among them the first records of the H. galilaeus from Pakistan and Taiwan. The enormous range of this
species, which now extends from the Middle East to Taiwan, is mapped. Unlike the
populations from other regions, the material of H. galilaeus from Taiwan
is brachypterous.
483:
On
the Staphylinidae of the Greek island Lesbos II, with supplementary notes on the
fauna of Samos and Chios (Coleoptera: Staphylinidae).
A field trip to the Greek island Lesbos conducted in spring 2016 yielded a
total of 1640 specimens belonging to 169 species of Staphylinidae. Nine new
species are described and illustrated: Myllaena lesbia Assing
sp.n., Aloconota (Aloconota) aegaea Assing
sp.n. (also recorded from Samos), and A. (A.) lesbia Assing
sp.n. of the Aleocharinae, Sunius potti Assing
sp.n. of the Paederinae, Bryaxis lesbius Brachat
sp.n., Bythinus simplicipalpis Brachat
sp.n., Protamaurops assingi Brachat
sp n., and Tychus lesbius Brachat
sp.n. of the Pselaphinae, and Stenichnus (Stenichnus) lesbius
Meybohm sp.n. of the Scydmaeninae.
The new species of Paederinae, Pselaphinae, and Scydmaeninae are most likely
endemic to Lesbos. A comprehensive list of the named and unnamed species
currently known from Lesbos is provided. Numerous described species are reported
from the island for the first time, eight of them represent first records from
Greece. The known staphylinid fauna of Lesbos currently includes 199 species,
184 named and 15 unnamed, unidentified, or tentatively identified. Nine (5 %) of
the named species and at least two of the unnamed Pselaphinae and Scydmaeninae
are hypothesized to be endemic to the island. Thus, despite greater overall
diversity and a generally similar composition, the fauna of Lesbos includes
fewer island endemics than that of the geographically close and significantly
smaller island Samos. As was to be expected, the fauna of Lesbos displays close
affiliations with that of the adjacent Turkish mainland. Several species reach
their westernmost distribution limit in this island (some of them also in Chios
and Samos). As many as six myrmecophilous species have been recorded from
Lesbos, five of them associated with Messor spp. and one with Cataglyphis
nodus (Brullι,
1833). In an appendix, Ocalea brachyptera Fagel,
1971 is reported from Chios, the first record of this species from Greece. Aloconota
(Aloconota) samia Assing
sp.n. (Samos) of the Aleocharinae and Stenichnus (Stenichnus) chius
Meybohm sp.n. (Chios) of the
Scydmaeninae are described and illustrated.
484:
A
revision of Geostiba of the West Palaearctic region. XXIII. On the Sibiota
species of the Caucasus region exclusive of Turkey (Coleoptera: Staphylinidae:
Aleocharinae).
Seven microphthalmous species of the subgenus Sibiota Casey, 1906, genus Geostiba
Thomson, 1858, from Georgia and
Armenia are described and illustrated: Geostiba (Sibiota) recta
nov.sp. (Georgia: Ratcha and Kvemo-Svanetia provinces); G. (S.) largata
nov.sp. (Georgia: Imereti and Ratcha); G. (S.) artifistula
nov.sp. (Georgia: Ratcha); G. (S.) unituber nov.sp. (North
Armenia: Pambaki range); G. (S.) pambakica nov.sp. (North
Armenia: Pambaki range); G. (S.) meghruica nov.sp. (South
Armenia: Meghru range); G. (S.) unicuneata nov.sp. (South
Armenia: environs of Jermuk). The type locality of G. carinicollis (Eppelsheim,
1878) is revised and G. medea Pace,
1996, previously a junior synonym of G. carinicollis, is revalidated.
Additional records of G. bituberculata (Eppelsheim,
1878) and G. kobrisensis Pace,
1996 are reported, and previous records of G. kobrisensis are rectified.
A key to the 17 Sibiota currently known from the Caucasus region
exclusive of Turkey, all of them microphthalmous and micropterous, is provided.
Their distributions are mapped.
485:
New
species of Oxypoda from Armenia and Georgia (Coleoptera: Staphylinidae:
Aleocharinae).
Six species of Oxypoda Mannerheim,
1830 from Armenia and Georgia are described and illustrated: Oxypoda
(Oxypoda) flexa nov.sp. (Armenia), O. (O.) articollis
nov.sp. (Georgia), O. (Deropoda) levipunctata nov.sp.
(Armenia), O. (Thliboptera) infissoides nov.sp. (Armenia), O.
(Bessopora) grandicristata nov.sp. (Armenia), and O. (Sphenoma)
subplicata nov.sp. (Armenia). Oxypoda recondita Kraatz,
1856, previously in the subgenus Baeoglena Thomson,
1867, is assigned to Bessopora Thomson,
1859.
486:
On
some species of the Quedius obliqueseriatus group, with notes on Q.
nivicola (Coleoptera: Staphylinidae: Staphylininae).
New data on several micropterous and locally endemic species distributed in the
Caucasus region and allied to Quedius obliqueseriatus Eppelsheim,
1889 are presented. Quedius nigrosuturalis nov.sp. (Northwest Georgia) is
described and illustrated. Quedius obliqueseriatus and Q. walteri Korge,
1971, both listed in Microsaurus Dejean,
1833 in the Palaearctic Catalogue, are assigned to the subgenus Raphirus Stephens,
1829. The distributions of Q. boluensis Korge,
1971, Q. smetanai Korge,
1971, and Q. walteri are mapped. Additional records are provided for Quedius
nivicola Kiesenwetter, 1858,
among them the first records from Turkey.
487:
Revision
of the Anaulacaspis species of the Palaearctic region (Coleoptera:
Staphylinidae: Aleocharinae).
Types
and additional material of the falagriine genus Anaulacaspis Ganglbauer,
1895 are revised. Disregarding one name of doubtful identity, 31 species are
recognized in the Palaearctic region, twelve of them new: Anaulacaspis
convexa sp. n. (Turkey); A. cristata sp. n. (North Pakistan); A.
eminens sp. n. (South Iran, Afghanistan); A. excisa sp. n. (South
Iran); A. flavomarginata sp. n. (Albania, Greece); A. gilva sp. n.
(North Pakistan); A. iberica sp. n. (Spain); A. pectinata sp. n.
(China: Yunnan); A. pseudonigra sp. n. (Turkey); A. reticulata sp.
n. (Turkey); A. sinuata sp. n. (Iran); A. truncata sp. n. (Iran).
One species, A. inexpectata (Fagel,
1969), is of doubtful generic assignment. The genus and the individual species
are (re-)described and illustrated. Twelve synonymies are proposed: Anaulacaspis
formosa (Rosenhauer,
1856) = A. cirrosa
(Fauvel,
1902), syn.
n., =
A. algirica
(Fagel,
1969), syn.
n., =
A. elkantarensis
(Fagel,
1969), syn.
n., =
A. macra
(Fagel,
1969), syn.
n., =
A. biskrensis
(Fagel,
1969), syn.
n.,
= A. andalusiaca
(Fagel,
1969), syn.
n.;
A. gratilla
(Erichson,
1839) =
A. lutzi
(Reitter,
1909), syn.
n.; A. naevula (Erichson, 1839) =
A. elegans
(Cameron,
1944), syn.
n., =
A. elegansides
Newton, 2015, syn.
n.; A. nigra (Gravenhorst,
1802) =
A. jonica
(Bernhauer,
1910), syn.
n., =
A. anatolica
(Fagel,
1969), syn.
n., =
A. corcyrana
(Fagel,
1969), syn.
n.
Lectotypes are designated for Aleochara nigra Gravenhorst, 1802, A. picea Gravenhorst, 1802, Falagriola lutzi Reitter,
1909, Falagria laeviuscula Eppelsheim,
1880, F. nigerrima
Bernhauer,
1908, and
F. beesoni Cameron, 1939. Based
on morphological characters, the species are assigned to six species groups.
Available data on the zoogeography and natural history of the Anaulacaspis
species recorded from the Palaearctic region are compiled. The distributions of
the genus as a whole and of the individual species are mapped. A catalogue and a
key to species are provided.
488:
On
the taxonomy and zoogeography of the species of the Quedius paradisianus
group (Coleoptera: Staphylinidae: Staphylininae).
The
distributions of four species of the Quedius paradisianus group Q.
paradisianus (Heer, 1839), Q.
italicus Gridelli, 1925, Q.
collaris Erichson, 1840, and Q.
rodopianus Coiffait, 1971
are revised, mapped, and discussed. While Q. italicus, Q. collaris,
and Q. rodopianus have discrete allopatric distributions, that of the
widespread and common Q. paradisianus significantly overlaps with the
ranges of the other three species. Several new country records are reported. Quedius
italicus, previously a subspecies of Q. collaris, is regarded as a
distinct species. Lectotypes are designated for Philonthus paradisianus Heer,
1839 and Quedius collaris Erichson,
1840. A key to species and illustrations of relevant diagnostic characters are
provided. Data on the natural history are compiled. The vertical distribution
and/or the phenology of three species are illustrated.
489:
On
the Lomechusini fauna of the East Palaearctic and Oriental regions, with a focus
on the genera Orphnebius and Amaurodera (Coleoptera:
Staphylinidae: Aleocharinae)..
Types
and additional material of 13 genera of Lomechusini from the Palaearctic and
Oriental regions are revised. In all, 28 species are (re-)described and/or
illustrated, 17 of them new: Orphnebius (Deroleptus) triacuminatus
sp. n. (Thailand) of the O. draco group; O. (D.)
gracilis sp. n. (South India) and O. (D.) protuberatus
sp. n. (South India) of the O. niger group; O. (D.)
pertortus sp. n. (India: West Bengal), O. (D.) migrus
sp. n. (India: Meghalaya), and O. (D.) reticulipennis sp.
n. (Indonesia: Java) of the O. siwalikensis group; Pheidologitonetes
bursata sp. n. (South India); Amaurodera bicarinata sp. n. (India:
Meghalaya); A. gilvicornis sp. n. (Thailand); A. latisulcata sp.
n. (Indonesia: Java); A. brevipes sp. n. (Indonesia: Sumatra); A.
migritheca sp. n. (Indonesia: Sumatra); A. longisetosa sp. n.
(Malaysia: Sabah); A. calicitheca sp. n. (Malaysia: Sabah); Drusilla
lativentris sp. n. (China: Yunnan); Rabdotodrusilla vara sp. n.
(India: Meghalaya); Witteia tensa sp. n. (China: Yunnan). The following
new combination and synonymy are established: Pheidologitonetes adesi (Pace,
1998), comb. n. (ex Zyrastilbus); Drusilla obliqua (Bernhauer,
1916) = D. palata Assing,
2015, syn. n. Zyrastilbus angkorensis Pace,
2004 is not congeneric with the type species of Zyrastilbus Cameron,
1939; its correct generic assignment (Drusilla?) is currently unknown. An updated catalogue of the 174 described representatives of Orphnebius
Motschulsky, 1858 in the East
Palaearctic and Oriental regions is provided; the species are grouped according
to a recently proposed intrageneric system. Amaurodera Fauvel,
1905 currently includes 53 described species distributed in the southern East
Palaearctic and the Oriental regions. Additional records of 46 named and three
unnamed species are reported, among them several new country records. The
distributions of two Himalayan Orphnebius species are mapped..
490:
On
Zyras sensu strictu in the East Palaearctic and Oriental regions, with a
focus on the faunas of the Himalaya, India, Stri Lanka, Thailand, and Sulawesi
(Coleoptera: Staphylinidae: Aleocharinae: Lomechusini).
The
species of the nominal subgenus of the lomechusine genus Zyras
Stephens,
1835 of the Himalaya, India, Sri Lanka, Thailand, and Sulawesi are revised.
Additional species from other parts of the East Palaearctic and Oriental regions
are addressed. In all, 40 species are described and/or illustrated, 14 of them
for the first time: Zyras
(Zyras)
ambulans
spec. nov. (Thailand), Z.
(Z.)
brevilobatus
spec. nov. (Thailand), Z.
(Z.)
densihirtus
spec. nov. (Sulawesi Utara), Z.
(Z.)
densissimus
spec. nov. (Sulawesi Utara), Z.
(Z.)
latilobatus
spec. nov. (South India), Z.
(Z.)
longilobatus
spec. nov. (India: Meghalaya), Z.
(Z.)
luteipes
spec. nov. (India: Meghalaya), Z.
(Z.)
morulus
spec. nov. (Nepal: Dhaulagiri, Annapurna), Z.
(Z.)
nigrihirtus
spec. nov. (Sulawesi Utara), Z.
(Z.)
parahirtus
spec. nov. (Borneo), Z.
(Z.)
parvicollis
spec. nov. (Thailand), Z.
(Z.)
russiceps
spec.
nov. (Thailand, Malaysia), Z.
(Z.)
titan
spec. nov. (Sulawesi Utara), and Z.
(Z.)
truncatus
spec. nov. (Nepal: Dhaulagiri). As many as 28 synonymies are proposed: Zyras
alternans
(Cameron, 1925) = Z. optimus
Cameron, 1939, syn. nov.; Z.
bartolozzii Pace,
2003 = Z. alboterminalis Pace,
2008, syn. nov.; Z. bettotanus Cameron,
1930 = Z. drescheri Cameron,
1939, syn. nov., = Z. atrapicalis Assing,
2016, syn. nov.; Z. brignolii (Pace,
1986) = Z. thainiger Pace,
2012, syn. nov.; Z. castaneus (Motschulsky,
1861) = Z. adulescens (Pace,
1987), syn. nov., =
Z. britannorum Pace,
1992, syn. nov., = Z. fratrumkadooriorum Pace,
1998, syn. nov., = Z. chumphonensis Pace,
2004, syn. nov., = Z. dibrugarhensis Pace,
2011, syn. nov.; Z. preangeranus Cameron,
1939 = Z. louwerensi Cameron,
1939, syn. nov., = Z. chinkiangensis
Bernhauer, 1939, syn. nov., = Z.
setosipennis Scheerpeltz, 1965,
syn. nov., = Z. alboantennatus Pace,
1986, syn. nov., = Z. sichuanorum Pace,
2012, syn. nov.; Z.
geminus (Kraatz,
1859) = Z. indicus Cameron,
1944, syn. nov., = Z. shiva Pace,
1992, syn. nov., = Z. manjushri
Pace, 1992, syn. nov., = Z. hongkongensis Pace,
1999, syn. nov., = Z. benenensis Pace, 2001, syn.
nov., = Z. parageminus Pace,
2010, syn. nov., = Z. neoparageminus
Hlavαč, Newton & Maruyama, 2011,
syn. nov., = Z. subgeminus Pace,
2012, syn. nov., = Z. articollis
Assing, 2016,
syn. nov.; Z. parageminus Pace,
1998 = Z. nameriensis Pace,
2011, syn. nov.; Z. pindarae (Champion, 1921) = Z. ruficauda Cameron,
1939, syn. nov.; Z. proximus Cameron,
1939 = Z. drugmandi Pace,
2004; syn. nov. Zyras novinversus
nom. n. is proposed for the preoccupied name Z.
inversus Pace,
2012, syn. nov. Lectotypes are
designated for Zyras exasperatus Schubert, 1908, Z. drescheri Cameron, 1939, Z. gratellus Cameron, 1939, Myrmedonia perforata Champion,
1921, and Hygroptera castanea Motschulsky,
1861. Remarkable cases of colour polymorphism and of sexual dimorphism
are discussed. A key to the Zyras sensu strictu species of the Himalaya,
India, and Sri Lanka, a key to the species recorded from Thailand, and an
updated catalogue of the Zyras sensu strictu species of the Palaearctic
and Oriental regions are provided. Additional records of numerous named and
several unnamed species are reported. The revised distributions of 37 species
are mapped. The subgenus is currently represented in the Palaearctic and
Oriental regions by a total of 124 described species. The countries with the
greatest diversity are China (46 named species), Malaysia (20), India (20),
Indonesia (18), Thailand (13), and Nepal (12). The subgenus is reported from
Sulawesi for the first time (five named and two unnamed species). Based on a
revision of type material, eleven species are not included in, or excluded from,
Zyras sensu strictu, and Zyras unicolor Cameron,
1939 is tentatively moved to Drusilla Leach,
1819..
491:
On
the Staphylinidae of the Greek island Kos, with an appendix on Carabidae and
additional records from other islands (Insecta: Coleoptera).
A
study of 567 specimens of Staphylinidae collected in the Greek island Kos in
April 2012 and December 2016 yielded 54 species, among them five first records
from Greece. One of them is Quedius curtidens Smetana, 1967, a species previously known only from Adana
province (Turkey) and also reported from Lebanon, Syria, and Israel for the
first time; its distribution is mapped. A checklist of the species recorded from
Kos is provided. The known staphylinid fauna is represented by 58 species and
shows strong affinities to the fauna of the close Turkish mainland. It is
primarily composed of widespread West Palaearctic (40 %), East Mediterranean (36
%), South Anatolian (11 %), or widespread Mediterranean (8 %) species. In an
appendix, records of 15 species are reported from Cyprus, Rhodos, Samos, and
Lesbos. Moreover, 29 species of Carabidae (177 specimens) are reported from Kos;
eight of these species are recorded from the Aegean Islands for the first time.
492:
On
Quedius coloratus Fauvel,
1875 and allied species, with an appendix on Quedius species collected in Greece
iwth subterranean pitfall traps, and a new synonymy (Coleoptera: Staphylinidae:
Staphylininae).
A revision of material previously identified as Quedius coloratus Fauvel,
1875 revealed that it is composed of at least five species: Q. (Raphirus)
coloratus (Middle East); Q. (R.) hellenicus nov.sp.
(Greece); Q. (R.) hebes nov.sp. (South Anatolia); Q. (R.)
spiculatus nov.sp. (North Anatolia); Q. (R.) carpathius
nov.sp. (Karpathos). These species, as well as one male possibly representing a
sixth species (Greece: Thessalνa) are described and illustrated. Available
evidence suggests that their habitats are subterranean and their distributions
are allo- or parapatric. All the named species are presumably capable of flight.
A key to species is provided. In an appendix, additional records of three Quedius
species collected in Greece with subterranean pitfall traps are reported. One of
these species is reported from Greece for the first time. The distributions of
the species allied to Q. coloratus, of Q. (Microsaurus) bernhaueri
Rambousek, 1915, and of Q. (Raphirus)
endogeus Assing,
2007 are mapped. Quedius endogeus, a species previously known only from
two localities, is widespread in Greece and evidently a true inhabitant of the
"Superficial Subterranean Habitat" (MSS stratum). Quedius
crnagoricus Coiffait, 1980,
nov.syn., is placed in synonymy with Q. paradisianus (Heer,
1839).
493:
A
revision of the Alevonota species of the Palaearctic region. II. A new
species from Cyprus and additional records (Coleoptera: Staphylinidae:
Aleocharinae).
Alevonota
flexa nov.sp.
(Cyprus) is described and illustrated. Additional records of four species are
reported, among them several new country records. Alevonota japonica (Cameron,
1933), a species with a remarkable sexual dimorphism of head shape and eye size,
is illustrated, based on new material from Hokkaido.
494:
Taxonomic
and faunistic notes on some West Palaearctic and Middle Asian Xantholinini, with
a revalidation and new synonymies (Coleoptera: Staphylinidae: Staphylininae).
Xantholinus
motschulskyi
Bordoni, 1999, previously a synonym
of X. reitteri Coiffait,
1966, is revalidated. The internal structures of the aedeagus of X.
motschulskyi, X. reitteri, and X. corallinus Reitter,
1901 are illustrated. A lectotype is designated for X. corallinus. Six
synonymies are proposed: Gyrohypnus punctulatus (Paykull,
1789) = Gyroyhypnus quadratus Stephens,
1833, nov.syn.; Stenistoderus cephalotes cephalotes (Kraatz,
1858) = S. cephalotes armeniacus (Coiffait,
1966), nov.syn.; Xantholinus linearis (Olivier,
1795) = X. styriacus Grimmer,
1841, nov.syn.; X. gridellii Coiffait,
1956 = X. tronqueti Bordoni,
2016, nov.syn.; X. khnzoriani Coiffait,
1966 = X. caucasicus Bordoni,
1975, nov.syn., = X. iablokoffi Coiffait,
1975, nov.syn. A neotype is designated for Gyroyhypnus quadratus Stephens,
1833. New records of 37 species of Xantholinini are reported. The revised
distributions of X. reitteri and X. motschulskyi are mapped.
495:
Two
new species and additional records of micropterous Paederus from China
and Taiwan (Coleoptera: Staphylinidae: Staphylininae).
Two
species of Paederus Fabricius,
1775 from China and Taiwan are described and illustrated: Paederus (Gnathopaederus)
furcillatus nov.sp. (China: Guizhou) and P. (Megalopaederus)
nigricrus nov.sp. (Taiwan: Taitung Hsien). Additional records of
eight species of Harpopaederus Scheerpeltz,
1957 (two species), Gnathopaederus Chapin,
1927 (one species), Megalopaederus Scheerpeltz,
1957 (three species), and of the P. biacutus group (two species) are
reported. The genus is currently represented in China and Taiwan by 51 and 25
species respectively.
496:
A
revision of Leptobium Casey.
VIII. A new species from Iran, a new synonymy, and additional records
(Coleoptera: Staphylinidae: Paederinae).
Leptobium
iranicum
nov.sp. (Iran: Yazd province) is described and illustrated. One synonymy is
proposed: Leptobium carinatum Assing,
2005 = nabozhenkoi Anlaş, 2012,
nov.syn. Additional records of 25 species and subspecies are reported. The genus
now includes a total of 67 species and two subspecies.
497:
Four
new species and additional records of Lathrobium from Nepal and China
(Coleoptera: Staphylinidae: Paederinae).
Four species of Lathrobium Gravenhorst,
1802 are described and illustrated: Lathrobium kongmaicum nov.sp.
(East Nepal: Kongma Danda), L. latilobatum nov.sp. (East Nepal: Kongma
Danda), and L. paulitortum nov.sp. (East Nepal: Solu Khumbu) of the L.
nepalense group, and L. piraticum nov.sp. (China: Sichuan: Wolong) of
the L. fissispinosum group. Additional records of three species from
China and three species from Nepal are reported, among them the first recent
records of L. nepalorientis Coiffait,
1984 and L. khumbuense Coiffait,
1982. The genus is now represented in the Palaearctic region by 595 species and
nine subspecies. The Himalayan fauna currently includes 76 described species,
all of them micropterous and locally endemic. As many as 220 named species have
been recorded from China.
498:
A
revision of Palaearctic Lobrathium. VI. Two new species from Armenia and
Taiwan, and additional records (Coleoptera: Staphylinidae: Paederinae).
Two species of Lobrathium Mulsant
& Rey, 1878 are described and illustrated: L.
ancoriferum nov.sp. (Armenia) and L.
biuncinatum nov.sp. (Taiwan). Additional records of 13 species are
presented. Lobrathium is now represented in the Palaearctic region
(including Myanmar and Vietnam) by 144 described species and one subspecies.
499:
A
revision of Palaearctic Nazeris IX. Three new species from Nepal and
additional records (Coleoptera: Staphylinidae: Paederinae).
Three species of Nazeris Fauvel,
1875 of the N. flavocaudatus group from East Nepal are described and
illustrated: Nazeris biapicalis nov.sp., N. hebes nov.sp., and N.
trapezilobus nov.sp. Additional records of three previously described
species are reported from East Nepal and Lebanon. The pronounced intraspecific
variation of external and sexual characters of N. ammonita (Saulcy,
1865) is discussed and illustrated. The genus now includes a total of 250
described species and seven subspecies. It is represented in the Himalaya
(exclusive of China) by 52 described species.
500:
A
revision of Neosclerus Cameron
VI. Thre new species from India, Thailand, and Vietnam (Coleoptera:
Staphylinidae: Paederinae).
Three species of Neosclerus
Cameron,
1924 are described and illustrated: N.
anguliceps
nov.sp. (South India), a species of uncertain affiliations, N.
concavus
nov.sp. (Thailand) of the N.
brevispinosus
group, and N.
vietnamensis
(Vietnam) of the N.
brevispinosus
group. The latter species represents the second record of the genus from
Vietnam. Including the new species, the
genus now includes 47 species distributed in the southern East Palaearctic and
Oriental regions.
501:
The
first species of Chinecallicerus from Sichuan, China (Coleoptera:
Aleocharinae: Geostibini).
Chinecallicerus
pinnatus
nov.sp. (China: Sichuan: Wolong), the first representative of the genus to be
recorded from outside Yunnan, is described, illustrated, and distinguished from
its congeners. An updated key to the species of Chinecallicerus Assing,
2004 is provided. The genus currently includes seven species, six of which are
representely solely by their respective holotypes.
502:
A
revision of types and additional material of Staphylinidae from the
Iablokoff-Khnzorian collection (Insecta: Coleoptera).
Types and additional material from the Iablokoff-Khnzorian collection are
revised. The following synonymies are proposed: Olophrum puncticolle Eppelsheim,
1880 = O. erevanicum Iablokoff-Khnzorian,
1964, nov.syn.; Astenus procerus (Gravenhorst,
1806) = Astenus silvicola Iablokoff-Khnzorian,
1961, nov.syn.; Myrmoecia reitteri (Eppelsheim,
1881) = Zyras argus Iablokoff-Khnzorian,
1960, nov.syn.; Aleochara (Xenochara) rambouseki Likovskύ,
1964 = A. addenda Likovskύ,
1981, nov.syn., = A. hamulata Assing,
2009, nov.syn. The holotypes of Rugilus armeniacus (Coiffait, 1970), Aleochara polychroma Iablokoff-Khnzorian,
1966, and A. nobilis Likovskύ,
1965 are illustrated. The latter is moved to the subgenus Xenochara Mulsant
& Rey, 1874 (ex Heterochara Mulsant
& Rey, 1874). Seven species of Staphylinidae are reported from
Armenia and one from Slovenia for the first time. The record of one species from
Armenia is based on a misentification.
503:
On
some Athetini from Armenia and adjacent regions (Coleoptera: Aleocharinae).
Nine species of Athetini from Armenia are described and illustrated, eight of
them for the first time: Atheta (Atheta) brevapicalis
nov.sp. (South Armenia); A. (A.) hamulata nov.sp. (South
Armenia); A. (Dimetrota) bispinosa nov.sp. (Armenia,
Georgia); A. (D.) senticollis nov.sp. (North Armenia); A.
(Oreostiba) altiviva Benick,
1974 (widespread in Armenia); A. (O.) semialba nov.sp.
(widespread in Armenia); A. (O.) abscisa nov.sp.
(widespread in Armenia); A. (O.) brevitheca nov.sp.
(widespread in Armenia); Liogluta
armeniaca nov.sp. (Central and South Armenia). Three synonymies are
proposed: Atheta hansseni Strand,
1943 = A. brachati Assing,
2013, nov.syn.; Liogluta granigera (Kiesenwetter,
1850) = L. trigemina (Eppelsheim,
1880), nov.syn.; L. funesta Eppelsheim,
1890 = L. nigrobusta Assing,
2016, nov.syn. A lectotype is designated for Homalota trigemina Eppelsheim,
1880.
504:
On
Stilicoderus VI. Two new species from South Vietnam and additional
records (Coleoptera: Paederinae).
Two species of Stilicoderus Sharp,
1889 from South Vietnam are described and illustrated: Stilicoderus
bisinuatus nov.sp. of the S. granulifrons group and S. harpago
nov.sp. of the S. discalis group. Additional records of five species are
reported, among them a new record from Bhutan. The genus now includes a total of
113 species.
506:
A
revision of Nazeris X. The first record of the genus from South Vietnam
and additional records from Thailand (Coleoptera: Staphylinidae: Paederinae).
Nazeris
raptus nov.sp. from South Vietnam is described and illustrated. This
record is of special zoogeographic significance, as it is situated nearly 6.5
degrees of latitude farther south than the previous southernmost record of the
genus as a whole, thus casting doubt on the previous classification of the
distribution of Nazeris Fauvel,
1873 as essentially South Palaearctic. Additional records of two species
from Thailand are reported. An updated map illustrating the distribution of the
genus as a whole is provided. Including the new species, Nazeris now
includes 255 described species and seven subspecies.
507:
Two
new species of Gnathymenus from Ecuador (Coleoptera: Staphylinidae:
Paederinae).
Two
species of the dolicaonine genus Gnathymenus Solier,
1849 from Ecuador are described and illustrated, Gnathymenus arboreus
nov.sp. and G. spinifer nov.sp., both of them macropterous and collected
by canopy fogging in the same locality in Pastaza province. The genus now
includes 77 species, 16 of which are known from Ecuador. Additional records of Gnathymenus
apterus Solier, 1849 are
reported from Chile.
508:
On
the micropterous Quedius (Raphirus) species with a punctate
scutellum of Turkey (Coleoptera: Staphylinidae: Staphylininae).
Four
micropterous species of Quedius Stephens,
1829, subgenus Raphirus Stephens,
1829, with a punctate scutellum are reported from North Turkey, one of them
unidentified. Quedius brevalatus nov.sp. (Armenia; Turkey: Rize) is
described and illustrated. Quedius omissus Coiffait, 1977, a species previously known only from the West
Caucasus, is reported from Turkey (Rize) for the first time. Three synonymies
are proposed: Quedius boops (Gravenhorst,
1802) = Philonthus boops tauricus Nordmann,
1837, nov.syn., = Quedius crius Tottenham,
1948, nov.syn. (previously a synonym of Q. reitteri Gridelli,
1925), = Quedius boops islandicus Fagel,
1960, nov.syn.
509:
A
mass dispersal event of Quedius hellenicus (Coleoptera: Staphylinidae:
Staphylininae).
In
spring 2017, a mass dispersal event of Quedius hellenicus Assing,
2017, the first observation of such a behaviour in the genus, was witnessed in
the Greek island Corfu. Additional evidence is presented suggesting that Q.
hellenicus inhabits a subterranean habitat and that this most likely also
applies to other representatives of the Q. coloratus group.
510:
On
the Leptusa fauna of the Caucasus region (Coleoptera: Staphylinidae:
Aleocharinae).
Five
species of Leptusa Kraatz,
1858 from Georgia and North Iran are described and illustrated: L. (Neopisalia)
substricta nov.sp. (Georgia: Shida Kartli); L. (N.) svanetica
nov.sp. (Georgia: Svaneti); L. (N.) migrituber nov.sp.
(Georgia: Svaneti); L. (N.) triangulata nov.sp. (Georgia:
Adjara); L. (N.) longalata nov.sp. (Iran: Mazandaran). The
genus is represented in the Caucasus region including Northeast Turkey (from
Ordu to the border with Georgia), Georgia, Armenia, Azerbaijan, and the Russian
part of the Greater Caucasus by 39 species in five subgenera; 28 species, two of
them of doubtful identity, belong to the subgenus Neopisalia Scheerpeltz,
1966. Three zoogeogeographically implausible or revised previous records are
rectified. Leptusa kaszabi Pace,
1981 is moved from the subgenus Heteroleptusa Pace,
1989 to Neopisalia. A zoogeographic study of the Leptusa fauna of
the Caucasus region revealed that locally endemic species are confined to the
moister western half of the region eastwards approximately to 43°30' eastern
longitude. Moreover, little is known about the fauna of Abkhazia, which probably
hosts an unknown number of unnamed species. A catalogue of the Leptusa
species of the Caucasus region and maps illustrating the currently known
distributions of 33 species are provided. Additional records of 19 described
species are reported, among them several new country records.
511:
On
the Geostiba fauna of Armenia (Coleoptera: Staphylinidae: Aleocharinae).
Three
microphthalmous species of the subgenus Sibiota Casey,
1906, genus Geostiba Thomson, 1858,
from Armenia are described and illustrated: Geostiba (Sibiota)
tigrani nov.sp. (North Armenia: mountains to the north of Sevan lake); G.
(S.) kalavanica nov.sp. (North Armenia: Kalavan mountain near the
northwestern tip of Sevan lake); G. (S.) immutata nov.sp.
(South Armenia: Barkushati range). Geostiba (Geostiba) sororcula
Assing, 2001, a species previously
known only from Northeast Turkey, is reported from Armenia for the first time.
Like its close relative, the widespread G. circellaris (Gravenhorst,
1806), this species is wing-dimorphic. At present, the known Geostiba
fauna is composed of nine described species in three subgenera; one or two
additional species are unnamed (males unavailable). Seven described and the
unnamed species, all of them microphthalmous, micropterous, and more or less
locally endemic, belong to the subgenus Sibiota; they are subject to
remarkable intraspecific variation of the male secondary sexual characters. A
key to species and distribution maps are provided.
512:
A
revision of Geostiba of the West Palaearctic region. XXV. New species
from Georgia and Greece, and additional records (Coleoptera: Staphylinidae:
Aleocharinae).
Four
species of Geostiba Thomson, 1858 from
Georgia and Greece are described and illustrated: Geostiba (Tropogastrosipalia)
xirosica nov.sp. (Greece: Evvoia); G. (T.) svanetica
nov.sp. (Georgia: Svaneti); G. (T.) gibberiventris nov.sp.
(Georgia: Mtskheta-Mtianeti); G. (Sibiota) granisuturalis
nov.sp. (Georgia: Svaneti). Additional records of seven species are reported.
The distributions of the eight species of the subgenus Tropogastrosipalia
Scheerpeltz, 1951 recorded from the
Caucasus region, those of the nine species of Sibiota Casey,
1906 known from the West and Central Greater Caucasus, and the distribution of G.
(Chondridiosipalia) cingulata (Eppelsheim,
1878) in the Caucasus region are mapped. Supplements to recently published keys
to the Geostiba fauna of the West Palaearctic region are provided.
513:
On
the Ischnosoma fauna of Georgia (Coleoptera: Staphylinidae: Tachyporinae).
Three
species of the Ischnosoma spelaeum group are described and illustrated,
all of them most likely with very restricted distributions: Ischnosoma acutum
spec. nov. (Georgia:
Svaneti, Racha); I. barbigerum spec. nov. (Georgia: Svaneti); I. molle
spec. nov. (Georgia:
Adjara). Type material of I. major (Luze,
1901) and I. caucasicum Kocian,
1997 is revised. Additional records of previously described species of Ischnosoma
Stephens,
1829 are reported from Georgia, partly also from other regions. Seven species of
the genus have reliably been recorded from Georgia, two species of the I.
pictum group and widespread, five of the I. spelaeum group and with
restricted distributions. The presence of an additional species of the I.
spelaeum group, I. caucasicum, requires confirmation. The
distributions of the species of the I. spelaeum group are revised and
mapped. Several previous literature records are zoogeographically implausible
and consequently probably erroneous.
514:
On
the Zyras sensu strictu in the East Palaearctic and Oriental regions III,
with a focus on the Southeast of Continental Asia and the Sunda Islands
(Coleoptera: Staphylinidae: Aleocharinae: Lomechusini).
Species
of the nominal subgenus of the lomechusine genus Zyras
Stephens,
1835 of the southeastern Oriental region are revised. Eighteen species are
(re-)described and/or illustrated, six of them for the first time: Zyras
(Zyras)
latibasalis
spec. nov. (Java, tentatively recorded also from Laos), Z.
(Z.)
rutrilobatus
spec. nov. (Laos), Z.
(Z.)
parvilobatus
spec. nov. (Laos), Z.
(Z.)
hirtiventris
spec. nov. (Laos, Vietnam), Z.
(Z.)
hlavaci
spec. nov. (Malaysia:
Pahang, Selangor), Z.
(Z.)
lunatus
spec. nov. (Malaysia: Selangor). Eight synonymies are proposed: Zyras
bryanti
Cameron,
1943 = Z. mortuorum Pace,
1990, syn. nov., = Z. paederinus Pace,
2008, syn. nov.; Zyras nigerrimus
Cameron,
1943 = Z. bartolozzii Pace,
2003, syn. nov., = Z. alboterminalis Pace,
2008, syn. nov.; Z. granulipennis Cameron,
1930 = Z. pervariolosus Pace,
2008, syn. nov.; Z. matangensis Cameron,
1943 = Z. daiaccorum Pace,
2008, syn. nov.; Z. montanus (Bernhauer,
1915) = Z. variolatus
Pace,
2003, syn. nov.;
Z. preangeranus
Cameron,
1939 = Z. quadriterminalis Pace,
2008,
syn. nov. One species is excluded from Zyras: Myrmedonota modiglianii
(Cameron,
1925), comb. nov. Lectotypes are designated for Zyras elegantulus Cameron,
1939, Z. semirufus Cameron,
1939, and Z. flavus Cameron,
1939. Additional
records of nine named species are reported. A key to the species recorded from
the southeastern Oriental region and an updated catalogue of the species of the
Palaearctic and Oriental regions are provided. The subgenus is currently
represented in the Palaearctic and Oriental regions by a total of 122 described
species. Twenty-two species have been recorded from Sunda Islands. The countries
with the greatest diversity are China (46 named species), India (20), Indonesia
(19), Malaysia (15), Thailand (14), Nepal (12), Laos (11), and Vietnam (8).
515:
A
revision of Sunius XVI. Two new species from Iran and Iraq (Coleoptera:
Staphylinidae: Paederinae).
Two
species of Sunius Stephens,
1829 are described and illustrated: S. reuteri spec. nov. (North Iraq),
the first record of the genus from Iraq, and S. concurvatus spec. nov.
(South Iran). Additional records of six previously described species are
reported from the West Palaearctic region. Updated distribution maps are
provided for S. khnzoriani (Coiffait,
1970) and S. fulgocephalus (Coiffait,
1970). The genus is currently represented in the Palaearctic region by 138
described species and two subspecies.
516:
On
the Staphylinidae of Crete III. The first records of endogean fauna (Coleoptera:
Staphylinidae: Leptotyphlinae, Aleocharinae).
Endogean
Staphylinidae were previously unknown from the Greek island Crete. During a
field trip conducted to East Crete in December and January 2017/2018 four
species of endogean Staphylinidae, three of the Leptotyphlinae and one of the
Aleocharinae, were collected using the soil-washing method. Two of the
leptotyphline species are described and illustrated: Kenotyphlus creticus
nov.sp. and Allotyphlus (Moreotyphlus) candicus nov.sp. The
third species of Leptotyphlinae belongs to the genus Gynotyphlus Coiffait, 1955 and remains undescribed for want of males.
Typhlocyptus pandellei Saulcy, 1878 (Aleocharinae)
is reported from Crete and Montenegro for the first time.
517:
The
third hypogean Domene species from Greece (Coleoptera: Staphylinidae:
Paederinae).
TDomene
(Domene)
vailatii
nov.sp. (Greece: Thesprotνa), the third hypogean representative of Domene
Fauvel, 1873 from Greece, is
described and illustrated. The species is apparently subject to remarkably
biased sex ratio; only one of the 22 type specimens is a male. The distributions
of the four hypogean Domene species known from Greece and Albania are
mapped.
518:
A
revision of Othiini. XX. Two new species of Othius, the first record of
the genus from Myanmar, and additional records (Coleoptera: Staphylinidae:
Staphylininae).
Othius
arater
nov.sp. (China: South Sichuan) and O. burmensis nov.sp. (Northeast
Myanmar), the first representative of the genus recorded from Myanmar, are
described and illustrated. New illustrations of the external and male sexual
characters of O. discrepans Assing,
1999 are provided. Additional records of ten species are reported. The
distributions of three Caucasian species are mapped. Othius Stephens,
1829 and the Othiini now include a total of 130 and 147 species and subspecies,
respectively.
519:
On
Hypnogyra laevissima (Reitter,
1898) (Coleoptera: Staphylinidae: Staphylininae: Xantholinini).
Based
on a revision of types and additional material, Hypnogyra laevissima (Reitter,
1898), a name recently placed in synonymy with the widespread H.
angularis (Ganglbauer, 1895), is revalidated. The external and male
sexual characters are illustrated and compared with those of H. angularis.
The known distribution of H. laevissima, which is confined to East
Azerbaijan and North Iran, is mapped.
520:
Six
new species of Chinecallicerus from China, with a new synonymy
(Coleoptera: Staphylinidae: Aleocharinae: Geostibini).
Six
species of
Chinecallicerus
Assing, 2004 are described and
illustrated:
Chinecallicerus
carinatus
nov.sp. (China: North Sichuan), C. orbiculatus nov.sp. (China: South
Sichuan), C. transversus nov.sp. (China: North Sichuan), C.
granulosissimus nov.sp. (China: East Qinghai), C. grandicollis
nov.sp. (China: Northwest Yunnan), and C. trituberculatus nov.sp. (China:
Northwest Yunnan). The following synonymy is proposed: Chinecallicerus
schuelkei Assing, 2004 = Aloconota
sinica Pace, 2011, nov.syn. An
updated key to the Chinecallicerus species is provided. Including the new
species, the genus now includes 13 species, eight of which have been recorded
from the Chinese province Yunnan, four from Sichuan, and one from Qinghai. All
the species are known only from their respective type localities, nine of them
are representely solely by their respective holotypes. The distributions of all
the species of the genus are mapped.
521:
On
the Nepalota fauna of China II. Two new synonymies and an additional
record (Coleoptera: Staphylinidae: Aleocharinae).
Two
synonymies are proposed: Nepalota cuneata Assing, 2015 = N. gibbera
Pace, 2017, nov.syn.; N.
tuberifera Assing, 2015 = N.
tengchongensis Pace, 2017,
nov.syn. A new record of N. chinensis Pace,
1998 from the Chinese province Jiangxi is provided.
522:
A
new Atheta species from the West Caucasus and a redescription of A.
brevapicalis (Coleoptera: Staphylinidae: Aleocharinae).
Atheta
("Alaobia") specicola nov.sp. (Russia: Krasnodarskiy
Kray), a cavernicolous species closely allied to A. trinotata (Kraatz,
1856), and A. brevapicalis Assing
& Vogel, 2017 are (re-)described and illustrated. Newly collected
material revealed that the type series of A. brevapicalis is composed of
two species. Additional records of A. brevapicalis, including the first
records from Iran, are reported; the distribution of the species is mapped.
523:
New
species and additional records of Cephalocousya and Drepasiagonusa
from China and Russia (Coleoptera: Staphylinidae: Aleocharinae: Oxypodini).
One
species of Cephalocousya Lohse,
1971 and four species of Drepasiagonusa Pace,
2012 are described and illustrated: Cephalocousya yunnanica nov.sp.
(China: North Yunnan), the first record of the genus from China; Drepasiagonusa
procera nov.sp. (China: North Yunnan); D. tibetica nov.sp. (China:
North Yunnan); D. angulata nov.sp. (China: Qinghai, Gansu); D.
feldmanni nov.sp. (Russia: East Siberia). The previously unknown aedeagus of
D. smetanai Pace,
2012, the type species and previously the sole representative of the genus, is
illustrated. Two additional Drepasiagonusa species from Shaanxi are
represented exclusively by females and remain unnamed. A key to the described
species of Drepasiagonusa is provided. The currently known distribution
of the genus is mapped. Cephalocousya nivicola is reported from
Kyrgyzstan for the first time.
524:
Six
new species of Trichoglossina from China (Coleoptera: Staphylinidae:
Aleocharinae: Oxypodini).
Six
species of Trichoglossina Pace,
1987
from China are described and illustrated: Trichoglossina aptera nov.sp.
(North Sichuan), T. alticola nov.sp. (North Sichuan), T. acutissima
nov.sp. (North Sichuan), T. volans nov.sp. (North Sichuan), T.
discolor nov.sp. (Shaanxi), and T. bifida (Northeast Yunnan). The
distributions of these species are mapped. The genus, which has been recorded
only from Nepal and China, now includes 39 species.
525:
Three
new species of Oxypoda from Spain, Armenia, and Ukraine, with notes on
the fauna of Armenia (Coleoptera: Staphylinidae: Aleocharinae).
Three
species of Oxypoda Mannerheim,
1830 from Spain, Armenia, and Ukraine are described and illustrated: Oxypoda
(Bessopora) parvioculata nov.sp. (Spain: Andalucνa: Sierra
Nevada), O. (Podoxya) verminata nov.sp. (Armenia), and O.
breviata nov.sp. (Ukraine). Additional records of 18 species are reported
from Armenia, 13 of them for the first time. The currently known Armenian Oxypoda
fauna is composed of at least 22 species, 20 of them named and 6 exclusive. The
currently known distributions of six species are mapped.
526:
On
the Aleochara subgenera Ceranota and Xenochara. IV. A
revision of types, a new species, and additional records (Coleoptera:
Staphylinidae: Aleocharinae).
Aleochara
(Ceranota) plicelytrata nov.sp. (Armenia, Georgia) of the A.
subtumida group is described and illustrated. Aleochara
(Xenochara)
citellorum
Kirshenblat,
1935 and
A.
(X.)
jacobsoni Kirshenblat,
1935 are
revised, redescribed, and illustrated; lectotypes are designated for both
species. They are assigned to the A. parvicornis group. Additional
records are reported for 13 species of Ceranota Stephens,
1839 and 18 species of Xenochara Mulsant
& Rey, 1874, among them 17 first records for Armenia (six), Georgia
(three), Ukraine (three), Russia (two), Lebanon (two), Greece (one), Iran (one),
and Kashmir (one). The distributions of six Ceranota and one Xenochara
species are mapped.
527:
On
the taxonomy and zoogeography of some West Palaearctic Quedius species,
with a focus on the East Mediterranean and the species allied to Quedius
umbrinus and Q. nivicola (Coleoptera: Staphylinidae: Staphylininae).
Based
on a revision of type material, as well as on studies of original descriptions
and intraspecific variation, 21 synonymies are proposed in the subgenus Raphirus
Stephens, 1829 of the genus Quedius
Stephens, 1829: Quedius
umbrinus Erichson, 1839 = Q.
cyanescens Mulsant & Rey, 1876,
nov.syn., = Q. bulgaricus Scheerpeltz,
1937, nov.syn., = Q. cyprusensis Last, 1955, nov.syn., = Q.
freyi Scheerpeltz, 1956,
nov.syn., = Q. maronitus Coiffait,
1963, nov.syn., = Q. gueorguievi Coiffait, 1967, nov.syn.; Quedius hermonensis Coiffait, 1963
= Q. coiffaitianus Fagel,
1968, nov.syn., = Q. rugosipennis Fagel, 1969, nov.syn.; Quedius
illyricus Wendeler, 1928 = Q.
paganettii Bernhauer, 1936, nov.syn.,
= Q. schipkanus Scheerpeltz,
1937, nov.syn., = Q.
pseudopyrenaeus Coiffait, 1967, nov.syn.;
Quedius nemoralis Baudi
di Selve, 1848 = Q. safaensis Fagel, 1968, nov.syn., = Q.
safaensis ormanus Fagel, 1971, nov.syn.,
= Q. nemoralis erinci Korge,
1971, nov.syn.; Quedius
limbatus (Heer, 1839) = Q.
scheerpeltzianus Fagel, 1968,
nov.syn.; Quedius suturalis
Kiesenwetter, 1845 = Q.
humeralis anatolicus Korge, 1964, nov.syn.,
= Q. troglophilus Coiffait,
1969, nov.syn.; Quedius job Coiffait, 1963 = Q.
lydus Fagel, 1968,
nov.syn.; Quedius humeralis Stephens,
1832 = Q. coxalis Kraatz,
1858, nov.syn., = Q.
atticus Coiffait, 1967, nov.syn.;
Quedius boops (Gravenhorst,
1802) = Q. haafi Scheerpeltz,
1956, nov.syn. Lectotypes are designated for Quedius cyanescens Mulsant
& Rey, 1876, Q. bulgaricus Scheerpeltz,
1937, Q. albanicus Bernhauer, 1926,
and Q. schipkanus Scheerpeltz,
1937. A neotype is designated for Q.
josue Saulcy, 1865. The male
sexual characters of several species are illustrated. Two species are
redescribed. The East Mediterranean species allied to Quedius
nivicola Kiesenwetter, 1858 are
revised. Their zoogeography and natural history are discussed. The currently
known distributions of ten species are mapped.
529:
A
revision of Medon. XI. Five new species, additional recortds, and the
first confirmed records from the Oriental region (Coleoptera: Staphylinidae:
Paederinae).
Five
species of the paederine genus Medon Stephens,
1833 from the East Palaearctic and Oriental regions are described and
illustrated: Medon arunicus nov.sp.
(East Nepal: Arun valley) and M.
acutapicalis nov.sp. (East Nepal: Theratum district) of the M.
apicalis group; M. sundaicus
nov.sp. (Indonesia: Java, Sulawesi) and M.
sabahicus nov.sp. (Malaysia: Sabah) of the M. ferrugineus group; M.
emeianus nov.sp. (China: Emei Shan) of the M. profundus group. Medon sundaicus
and M. sabahicus represent the
first confirmed records of the genus from the Oriental region. Additional
records of 32 species and subspecies are reported, among them several new
country records. Medon
is now represented in the Palaearctic region by five species groups and
86 confirmed species, 23 of which are distributed in the East Palaearctic
region.
530:
On
the Staphylinidae of the Greek island Corfu (Insecta: Coleoptera).
A
study of nearly 10,000 specimens of Staphylinidae collected in the Ionian island
Corfu, Greece, in late spring 2017 yielded more than 233 species. Additional,
previously unpublished records of 66 named species are reported from the island.
Two species are described and illustrated: Borboropora corcyrana Assing
spec. nov. of the Aleocharinae and Ocypus corcyranus Assing
spec. nov. of the Staphylininae. As many as 118 named species are reported from
Corfu for the first time, 21 of these species represent first records from
Greece. One name is revalidated and six names are synonymized: Euplectus
jonicus Meggiolaro, 1966
(revalidated) = E. jonicus
corcyreus Meggiolaro, 1966,
syn. nov.; Mycetoporus punctipennis Scriba,
1868 = M. insulanus Luze, 1901,
syn. nov.; Anotylus tetracarinatus (Block,
1799) = A. corcyranus (Coiffait,
1968), syn. nov.; Bledius corniger Rosenhauer,
1856 = B. bubalus Gistel,
1857, syn. nov.; Paederus littoralis Gravenhorst,
1802 = P. pelikani Reitter,
1884, syn. nov.; Leptacinus batychrus (Gyllenhal,
1827) = Phacophallus corcyranus Bordoni,
2017, syn. nov. Including reliable previous literature records and the new
records reported in the present paper, 446 named species (plus additional
unnamed species) are currently known from Corfu. Thus, the known fauna of this
island is significantly more diverse than those of other East Mediterranean
islands, including the much larger Cyprus. A checklist of the Staphylinidae
fauna of Corfu is provided. Although at present 18 species and subspecies have
been recorded exclusively from Corfu, most of them are unlikely to represent
island endemics; three of these species are of doubtful taxonomic status.
531:
On
the subterranean Staphylinidae fauna of South Spain (Coleoptera).
Staphylinidae
from soil samples collected in South Spain, all of them obtained by soil
washing, were examined. Seven species belonging to three subfamilies are
described and illustrated: Alevonota alboculata sp. n. (Murcia) of the
Aleocharinae, Lusitanopsis lencinai sp. n. (Murcia) of the Osoriinae, and
Entomoculia (Stenotyphlus) solanae sp. n. (Castilla-La
Mancha: Albacete), Paratyphlus deformis sp. n. (Murcia), P. magnispina
sp. n. (Castilla-La Mancha: Ciudad Real), P. lencinai sp. n. (Murcia),
and P. carmeloi sp. n. (Murcia) of the Leptotyphlinae. Paraleptusa
anophthalma (Eppelsheim,
1884) is recorded for the first time since the original description; its habitus
and the primary sexual characters are illustrated. An overview of the
subterranean Staphylinidae fauna previously known from South Spain is provided.
The distributions of three genera in (the south of) the Iberian Peninsula are
mapped. New records of some additional species are reported.
532:
On
some species of Apatetica Westwood
(Coleoptera: Staphylinidae: Apateticinae).
Eight
species of Apatetica
Westwood,
1848 are (re-)described and illustrated, six of them for the first time: Apatetica
viridipennis Fauvel, 1895
(North India); A.
glaucipennis spec. nov. (Nepal);
A. confusa spec. nov. (Myanmar;
previously confounded with A. viridipennis); A. aspera spec. nov.
(China: Sichuan); A. laevicollis Fauvel, 1904 (Vietnam); A. glabra spec. nov. (China:
Yunnan); A. gibba spec. nov. (China: Gansu); A. curtipennis spec.
nov. (China:
Yunnan). A lectotype is designated for Apatetica laevicollis. The first
confirmed record of A. laevicollis since the original description is
reported. The genus currently includes 25 described species distributed in the
southern East Palaearctic and Oriental regions.
533:
Five
new species of Lathrobium from Nepal and China (Coleoptera:
Staphylinidae: Paederinae).
Five
species of the Holarctic genus Lathrobium Gravenhorst,
1802 are described and illustrated: L.
adsurgens spec. nov. (Northwest Nepal: Gurans Himal) of the L.
deuvei group, L. guransicum
spec. nov. (Northwest Nepal: Gurans Himal) of the L.
emodense group, L. latius spec. nov.
(Northwest Nepal: Karnali) of the L.
jumlense group, L.
egens spec. nov. (China: Sichuan), a close relative of L.
diffissum Assing,
2013, and L. bitaleatum spec. nov.
(China: Sichuan) of the L.
bibaculatum group. Lathrobium is currently represented in the Palaearctic region by as many as 612
described species, 222 of which have been recorded from China and 79 from the
Himalaya.
534:
Revision
of the Cousya species of the West Palaearctic Region (Coleoptera: Staphylinidae:
Aleocharinae).
Species
of the genus Cousya Mulsant & Rey, 1875 and five previously unrevised
species of Ocyusa Kraatz, 1856 of the West Palaearctic region including
Middle Asia are revised. In all, 19 species of Cousya are recognized,
with one additional species of which no material was available of doubtful
identity and generic affiliations. Three or four additional species remain
unnamed for want of males. Only two species, O. maura (Erichson, 1837)
and O. picina (Aubι, 1850), remain in Ocyusa. Four species are
newly described: Cousya acris n. sp. (South Greece, South Turkey,
Cyprus, Lebanon); C. struyvei n. sp. (South Spain); C. pauli
n. sp. (Southwest Turkey); C. sufflata n. sp. (Greece:
South Pelopσnnisos). Redescriptions are provided for Cousya and ten of
its species. All the Cousya species and most of the species moved to
other genera are illustrated. Eleven synonymies are proposed: Cousya bicolor
(Bernhauer, 1900) = C. mirabilis Assing, 2011, n. syn.; C.
defecta (Mulsant & Rey, 1875) = C. humicola Fagel, 1965, n.
syn.; C. longitarsis (Thomson, 1867) = C. rugipennis (J.
Sahlberg, 1890) n. syn. (previously a synonym of Oxypoda funebris
Kraatz, 1856), = C. peezi Scheerpeltz, 1957, n. syn.; C.
nigrata (Fairmaire & Laboulbθne, 1856) = C. cephallenica
(Scheerpeltz, 1931), n. syn., = C. nitidiventris Fagel, 1958, n.
syn., = C. lakloukensis Fagel, 1965, n. syn.; C. schuelkei
Assing, 2007 = C. planicollis Assing, 2011, n. syn.; C.
bimaculata (Fauvel, 1899) = C. vaulogeri (Bernhauer, 1936), n.
syn.; Oxypoda flavicornis Kraatz, 1856 = Ocyusa beieri
Scheerpeltz, 1931, n. syn.; Ocyusa pellax Peyerimhoff, 1919 = Oxypoda
argus Normand, 1935, n. syn. Six new binomina are established: Cousya
dissoluta (Eppelsheim, 1888), n. comb. (ex
Ocyusa); C. praecox (Eppelsheim, 1888), n. comb. (ex Ocyusa);
Oxypoda picta Mulsant & Rey, 1875, n. comb. (ex Cousya);
Oxypoda pellax (Peyerimhoff, 1919), n. comb. (ex Ocyusa); Oxypoda
heydeni (Eppelsheim, 1879), n. comb. (ex Ocyusa); Tectusa
uhligi (Pace, 1987), n. comb. (ex
Cousya). The material labelled as types of Cousya uhligi is
composed of three species, all of them belonging to Tectusa Bernhauer,
1899. One of them is Tectusa pirinica n. sp. (Bulgaria:
Pirin Planina). Four
Bulgarian species of Tectusa, including T. pirinica, are
illustrated. Ocyusa apicalis Normand, 1935 is not congeneric
with the type species of Ocyusa and treated as Oxypodina incertae sedis.
A neotype is designated for Ocyusa defecta Mulsant & Rey, 1875.
Lectotypes are designated for Cousya peezi Scheerpeltz, 1957, Ocyusa
eppelsheimi Bernhauer, 1902, O. araxis Bernhauer, 1902, O. bicolor
Bernhauer, 1900, Homalota dissoluta Eppelsheim, 1888, H. praecox
Eppelsheim, 1888, and Ocyusa ferdinandicoburgi Rambousek, 1909. A
catalogue and a key to the Cousya species of the West Palaearctic region
are provided. The species of Cousya are assigned to two species groups,
the C. nigrata group (eleven species) and the C. crocea group
(eight species). While the species of the C. nigrata group are fully
winged and widespread, those of the C. crocea group are partly incapable
of flight and have restricted distributions; six of them have been recorded only
from their respective type localities. The distributions of all the Cousya
species are mapped. Available evidence suggests that at least the species of the
C. nigrata group reproduce in autumn (most species in a subterranean
habitat), have their preimaginal development in winter and early spring, emerge
from the pupa and disperse in spring, and spend the warmer seasons either in
estivation or in a subterranean habitat. A biased sex ratio (males rarer than
females) was observed in two species. It is particularly pronounced in C.
nigrata: only 15 % of the sexed specimens are males. Not a single male was
found in the north of its range, suggesting that the species is parthenogenetic
in this region.
535:
An
enigmatic new genus and species of Anthophagini from Georgia (Coleoptera:
Staphylinidae: Omaliinae).
Caucanthobium
nov.gen.
mirandum nov.sp., an enigmatic new genus and species from Southwest
Georgia, is described, illustrated, and distinguished from Anthobium Leach,
1819 and Deinopteroloma Jansson,
1947. Its currently known distribution is mapped.
536:
A
revision of Palaearctic and Oriental Scymbalium and Micrillus III.
New species, new combinations, and additional records (Coleoptera:
Staphylinidae: Paederinae).
Six
species of Scymbalium Erichson,
1839 and Micrillus Raffray, 1873
are described and illustrated: Scymbalium rossii nov.sp. (Cambodia), Micrillus
virgatus nov.sp. (Cambodia, South Thailand) of the newly established M.
badius group, M. hamatus nov.sp. (South Morocco) of the M.
testaceus group, M. kongi nov.sp. (Cambodia) of the M. suturalis
group, and M. coloratus nov.sp. (Cambodia) and M. rougemonti nov.sp.
(Sri Lanka) of the newly established M. coloratus group. Based on a
re-assessment of the inter- and intrageneric phylogenetic affiliations of Scymbalium
and Micrillus, two species are moved from Scymbalium to Micrillus:
Micrillus badius (Motschulsky,
1858), nov.comb.; Micrillus nepalensis (Assing,
2013), nov.comb. Additional records of one species of Scymbalium
and of ten species (one of them undescribed) of Micrillus are reported,
among them five first records from Cambodia (3), Syria (1), and Indonesia (1).
537:
A
revision of Pinobius. II. Three new species from Cambodia and Indonesia,
and additional records (Coleoptera: Staphylinidae: Paederinae).
Three
species of the dolicaonine genus Pinobius Macleay, 1871 are described and illustrated: Pinobius
acutus nov.sp. (Cambodia) and P. rossii nov.sp. (Cambodia) of the P.
indicus group, and P. forcipifer nov.sp. (Indonesia: Kalimantan
Timur) of the P. major group. Additional records of six previously
described species are reported, among them a new record from Cambodia. The genus
now includes a total of 38 described species, 30 of which are represented in the
southern Palaearctic, Oriental, and Australian regions.
538:
A
revision of Palaearctic and Oriental Pseudolathra. V. Two new species
from Cambodia and Thailand, and additional records (Coleoptera: Staphylinidae:
Paederinae).
SPseudolathra
armigera
nov.sp. (Cambodia) and P. aviformis nov.sp. (North Thailand) are
described and illustrated. One synonymy is proposed: Pseudolathra fissa Assing,
2013 = P. ceylonica Rougemont,
2015, nov.syn. Additional records of seven previously described species are
reported, among them several new country records. The genus is reported from
Cambodia (four species) and Bhutan for the first time and currently represented
in the East Palaearctic and Oriental regions by 40 described species. A
catalogue of these species is provided.
539:
On
some Myllaena species in the East Mediterranean and Caucasus regions
(Coleoptera: Staphylinidae: Aleocharinae).
Two
species of previously uncertain status of Myllaena Erichson,
1837 are revised, redescribed, and illustrated: M. graeca Kraatz,
1858 (Greece, Turkey) and M. caucasica Eppelsheim,
1880 (Caucasus region). A neotype and a lectotype are designated for M.
graeca and M. caucasica, respectively. Three species are described
and illustrated: M. mutabilis nov.sp. (Greece: Corfu), M. cretica
nov.sp. (Greece: Crete), and M. ambulans nov.sp. (Armenia). Eight first
records of Myllaena species from Armenia (3), Nagorno-Karabakh (2),
Russia (1), Azerbaijan (1), and Iran (1) are reported. The currently known
distributions of M. graeca, M. caucasica, and of the three newly
described species are mapped.
540:
A
revision of Geostiba of the West Palaearctic region. XXVI. New species
and additional records, primarily from the Caucasus region (Coleoptera:
Staphylinidae: Aleocharinae).
Seven
species of Geostiba Thomson,
1858 from the Caucasus region and Ukraine are described and illustrated: Geostiba
(Tropogastrosipalia) reducta nov.sp. (Russia: West Caucasus); G.
(T.) defecta nov.sp. (Russia: West Caucasus); G. (Sibiota)
breviflagellata nov.sp. (Georgia: Imereti); G. (S.) nigrohortensia
nov.sp. (Nagorno-Karabakh); G. (S.) deliqua nov.sp. (South
Armenia); G. (Sipalotricha) marmotae nov.sp. (Ukraine); G.
(S.) manca nov.sp. (Georgia: Imereti). The primary sexual
characters of several previously described species are illustrated. Geostiba
cingulata (Eppelsheim,
1878), previously in the subgenus Chondridiosipalia Scheerpeltz,
1951 is moved to the subgenus Sipalotricha Scheerpeltz,
1951. Additional records of eight Caucasian species are reported. The currently
known distributions of 15 Caucasian species of the subgenera Tropogastrosipalia
Scheerpeltz, 1951, Sibiota Casey,
1906, and Sipalotricha are mapped..
541:
A
new genus of Oxypodini from China (Coleoptera: Staphylinidae: Aleocharinae).
Notocousya
nov.gen. quadriceps nov.sp. (China: Yunnan) of the Oxypodini, subtribe
Oxypodina, is described and illustrated. The new genus currently includes only
the type species N. quadriceps and is characterized by a distinctive
morphology of the head, including the mouthparts, and of the pronotum. The
absence of previous records suggests a cryptic reproduction habitat.
542:
Revision
of the Baeoglena species of the West Palaearctic Region (Coleoptera:
Staphylinidae: Aleocharinae).
The
subgenus Baeoglena Thomson,
1867 of the speciose aleocharine genus Oxypoda Mannerheim, 1830 has been subject to considerable taxonomic
confusion rendering a reliable identification of material from regions other
than Central Europe and the Canary Islands virtually impossible. Based on a
revision of abundant material from various major public and private collections,
seven species are distributed in the West Palaearctic region exclusive of the
Canary Islands. Two new species are described and illustrated: Oxypoda (Baeoglena)
rectacia spec. nov (East Mediterranean, from South Greece to the Middle
East) and O. (B.) derecta spec. nov. (West Caucasus,
Northeast Anatolia). Diagnoses and illustrations of the genitalia are provided
for the remaining five species. The following synonymies are established: Oxypoda
nova Bernhauer, 1902 = O.
giachinoi Pace, 2001, syn.
nov.; O. hispanica Fagel,
1958 and its replacement name O. inexpectata Fagel, 1965 are removed from synonymy with O. praecox Erichson,
1839 and synonymized with O. fusina Mulsant
& Rey, 1875. Oxypoda recondita Kraatz,
1856 and O. dalmatina Bernhauer,
1905 are excluded from Baeoglena and moved to the subgenus Bessopora
Thomson, 1859. Lectotypes are
designated for Oxypoda nova Bernhauer,
1902, O. caucasica Bernhauer,
1902, and O. kuehnelti Scheerpeltz,
1963. An outline of the taxonomic history, a diagnosis of the subgenus, a
checklist, and a key to species are provided. A zoogeographic analysis revealed
some remarkably discontinuous distributions and parapatric distribution patterns
which are plausibly explained only with interspecific competition among Baeoglena
species. The distributions of the subgenus as a whole and of the individual
species in the West Palaearctic region are mapped.
543:
On
the Lomechusini fauna of the Palaearctic and Oriental regions. XXVI. New
species, a new synonymy, and additional records (Coleoptera: Staphylinidae:
Aleocharinae).
Types
and additional material of twelve genera of Lomechusini from the Palaearctic and
Oriental regions are revised. In all, 24 species are described and/or
illustrated, 21 of them new: Pella feldmanni spec. nov. (China: Sichuan)
of the P. cognata group; Orphnebius fortesetosus spec. nov.
(China: Sichuan) and O. taurus spec. nov. (Cambodia) of the O. hauseri
group; Orphnebius (Deroleptus) morulus spec. nov. (South
Vietnam), O. (D.) globulifer spec. nov. (Malaysia: Pahang),
O. (D.) elevatus spec. nov. (Laos), and O. (D.)
moruliflavus spec. nov. (Malaysia: Sabah) of the O. siwalikensis
group; Keratodegnathus cornutus spec. nov. (Malaysia:
Pahang); Amaurodera schmidti spec. nov. (East
Nepal) of the A. silvana group; A. brevipennis spec. nov. (Malaysia:
Sabah); A. atra spec. nov. (Philippines: Mindanao); Drusilla bispinosa
spec. nov. (Malaysia: Pahang); D. shavrini spec. nov. (Philippines:
Mindanao); D. penicillata spec. nov. (Philippines: Mindanao); D.
spiniventris spec. nov. (Philippines: Mindanao); D. breviuter spec.
nov. (Philippines: Mindanao); Pheidologitonetes biplicatus spec. nov. (China:
Yunnan); Zyras (Zyras) conlectus spec. nov. (Myanmar);
Z. (Z.) hetzeli spec. nov. (Taiwan);
Z. (Z.) reductus spec. nov. (Philippines: Mindanao); Z. (Z.)
shavrini spec. nov. (Philippines: Mindanao). The following synonymy is
established: Drusilla obliqua (Bernhauer,
1916) = D. zyrasoides Dvořαk,
1988,
syn. nov. Additional records of 49 named species are reported, among them twelve
first records from Greece (1), Lebanon (1), Israel (1), Jordan (1), Iran (1),
Nepal (1), China (1), Taiwan (1), Laos (1), Cambodia (2), and Borneo (1). The
distribution of Myrmoecia urartu (Iablokoff-Khnzorian, 1962) is mapped.
544:
The
Staphylinidae of Armenia and Nagorno-Karabakh (Coleoptera).
The
previously largely neglected and poorly known staphylinid faunas of Armenia and
Nagorno-Karabakh are addressed. Based on a study of more than 31,000
Staphylinidae recently collected in various habitats and using different
methods, and on a critical evaluation of previous literature records, a
checklist of the faunas of Armenia and Nagorno-Karabakh is compiled. The fauna
of Armenia currently includes 675, that of Nagorno-Karabakh 198 named species.
Nevertheless, it is concluded that the species inventory of both regions,
especially that of Nagorno-Karabakh, is still far from complete. As many as 262
and 183 species are reported from Armenia and Nagorno-Karabakh, respectively,
for the first time. A list of 99 species erroneously or doubtfully recorded from
the study region is provided. A comparison with the species number and
systematic composition of the faunas of other Caucasian countries and regions
revealed that (a) their known diversities are significantly lower than should be
expected and (b) a remarkably high proportion (nearly 40 %) of Aleocharinae in
the faunas of Armenia and Nagorno-Karabakh, with the genus Atheta Thomson,
1858 alone accounting for approximately 10 % of the total diversity in Armenia.
The faunas of Armenia and Nagorno-Karabakh are primarily composed of widespread
species, many of them Caspian (Caucasian) and Iranian elements, and remarkably
few endemics. Only 23 species, thirteen of the Aleocharinae (most of them
belonging to the genus Geostiba Thomson,
1858), four of the Pselaphinae, five of Scydmaeninae, and one of Staphylininae
are classified as regional endemics, and one species of Aleocharinae from the
peak region of Mount Khustup is classified as a local endemic. The distribution
of one addititional species is confined to South Armenia and adjacent parts of
North Iran. A number of species is currently known only from Armenia, but of
doubtful taxonomic status or unlikely to represent endemics. Records of some
species in Armenia and/or Nagorno-Karabakh revealed some remarkably
discontinuous distributions with gaps of up to approximately 2,800 km; three of
these distributions are mapped. Fourteen species are newly described: Omalium
kociani Zanetti spec. nov.
(Armenia: Jermuk) of the Omaliinae, Proteinus baculatus Assing
spec. nov. (Armenia; Northeast Turkey) of the Proteininae, Bryaxis armeniacus
Brachat spec. nov. (Armenia: NW Hrazdan) and B. meghruicus
Brachat spec nov. (South Armenia:
Meghru range) of the Pselaphinae, Atheta (Paralpinia) meghruica
Assing spec. nov. (South Armenia:
Meghru range), Bellatheta khustupica Assing
spec. nov. (South Armenia: Mount Khustup), Calodera alticola Assing
spec. nov. (Armenia: Mount Karkar), and Tachyusa unguis Assing
spec. nov. (South Armenia) of the Aleocharinae, Anotylus hamatoides Schόlke
spec. nov. (Armenia) of the Oxytelinae, Euconnus (Tetramelus) longilaminatus
Meybohm spec. nov. (North Armenia),
E. (T.) tavushus Meybohm
spec. nov. (North Armenia), E. (T.) karabakhus Meybohm
spec. nov. (Nagorno-Karabakh), Neuraphes (Paraphes) gomarantsus
Meybohm spec. nov. (South Armenia:
Meghru range), and N. (P.) syunikus Meybohm
spec. nov. (South Armenia) of the Scydmaeninae. Eight synonymies and one
revalidation are established: Dialycera minuta Luze,
1906 = Phyllodrepa armena Iablokoff-Khnzorian,
1959, syn. nov.; Mycetoporus
silvaticus Iablokoff-Khnzorian,
1962 = M. dispersus Schόlke &
Kocian, 2000, syn. nov.; Aleochara subtumida (Hochhuth,
1859) = Aleochara khnzoriani Amiryan,
1999, syn. nov.; Platystethus cephalotes Eppelsheim,
1878, revalidated (previously synonym of P. laevis Mδrkel &
Kiesenwetter, 1848) = P. oblongopunctatus Roubal, 1911, syn. nov.; Euconnus lalvarensis Iablokoff-Khnzorian,
1964 = Euconnus pseudorobustus Franz,
1986, syn. nov.; Astenus rufopacus Reitter,
1909 = A. baali Coiffait,
1960, syn. nov.; Heterothops dissimilis (Gravenhorst,
1802) = H. armeniacus Coiffait,
1977, syn. nov.; Heterothops praevius Erichson, 1839 = Heterothops montanus Iablokoff-Khnzorian,
1966, syn. nov.
545:
A
revision of Palaearctic and Oriental Rugilus. VI. Two new species from
China, a new synonymy, and additional records (Coleoptera: Staphylinidae:
Paederinae).
Two
species of Rugilus Leach,
1819 from China are described and illustrated: R. (Rugilus) reuteri
nov.sp. (North Sichuan) and R. (R.) gryps nov.sp. (West
Sichuan: Gongga Shan). Additional records of 27 previously described species, 18
of the nominal subgenus, seven of Eurystilicus Fagel,
1853, and two without subgeneric assignment, are reported from the Palaearctic
and Oriental regions, among them six new country records from Pakistan (3),
Lebanon (1), India (1), and Cambodia (1). A new synonymy is proposed: Rugilus
arabs (Saulcy, 1865) = R.
couloni (Drugmand, 1989),
nov.syn. The preoccupied name Rugilus rugossimus Assing, 2015 is replaced with R. asperrimus nov.nom. Stilicus
ovicollis MacLeay, 1873, a
species erroneously transferred to Rugilus recently, is reassigned to Scopaeus
Erichson, 1839. New illustrations
of the male sexual characters of R. prodoni (Coiffait, 1982), R. bagmaticola Rougemont,
1998, and R. curvatus Assing,
2013 are provided. The distributions of the species of the R. malaisei
group are mapped. Including the new species, Rugilus is now represented
in the Palaearctic and Oriental regions by 102 species and one subspecies; 35
species are currently known from China.
546:
A
revision of Palaearctic and Oriental Scymbalium and Micrillus IV.
The fauna of Cambodia (Coleoptera: Staphylinidae: Paederinae).
Records
of one species of Scymbalium Erichson,
1839 and seven species of Micrillus Raffray, 1873 are reported from Cambodia. Three of them are
described and illustrated: Micrillus rossianus nov.sp. of the M.
badius group, M. variceps nov.sp. of the newly established M.
variceps group, and M. bispinosus nov.sp. of the M. coloratus
group. Including these additions, the Cambodian fauna is currently composed of
one Scymbalium and nine Micrillus species and thus more diverse
than that of any other country in the Oriental region.
547:
Three
new species and additional records of Oedichirus (Coleoptera:
Staphylinidae: Paederinae).
Three
species of Oedichirus Erichson,
1839 are described and illustrated: O. pauli nov.sp. (North Vietnam), O.
puicus nov.sp. (North Thailand), and O. russipennis nov.sp.
(Indonesia: Papua: Yapen). A lectotype is designated for O. terminatus Erichson,
1843, a species described from Angola and subsequently reported also from the
Mediterranean region. The sexual characters of O. terminatus are
illustrated. Additional records of three species are provided.
548:
On
the taxonomy of some West Palaearctic Quedius species, with descriptions
of new species and new synonymies (Coleoptera: Staphylinidae: Staphylininae).
Three
species of Quedius Stephens, 1829
are described and illustrated, two of them new: Quedius (Raphirus)
iridicolor Quedenfeldt,
1882, Q. (Microsaurus) morulus nov.sp. (Iran), and Q.
(M.) limans nov.sp. (Israel). Three synonymies are established: Quedius
umbrinus Erichson, 1839 = Q.
kuboni tourač, 1998, nov.syn.; Q. cohaesus Eppelsheim,
1888 = Q. pseudonigriceps Reitter,
1909, nov.syn.; Q. cruentus (Olivier,
1795) = Q. xanthurus Iablokoff-Khnzorian, 1961, nov.syn. Diagnoses distinguishing Q.
iridicolor and Q. nigriceps Kraatz,
1857, as well as Q. (Microsaurus) invreae Gridelli,
1925 and Q. (M.) puncticollis (Thomson,
1867) are provided. Previous erroneous records of Q. puncticollis are
rectified. Quedius iridicolor is re-assigned to the subgenus Raphirus
Stephens, 1829. The currently known
distribution of Q. iridicolor is mapped.
549:
A
revision of the Alevonota species of the Palaearctic region. III. Two new
species from Taiwan, a new combination, a new synonymy, and additional records
(Coleoptera: Staphylinidae: Aleocharinae).
Alevonota
calliceroides nov.sp.
and A. hetzeli nov.sp., both of them from the same locality in Hualien
Hsien, Taiwan, are described and illustrated. Homalota hepatica Erichson,
1839, the type species of Enalodroma Thomson,
1859, is transferred to Alevonota Thomson,
1858, rendering Enalodroma a junior synonym of Alevonota and
yielding the binomen Alevonota hepatica (Erichson,
1839), nov.comb. Data on the natural history of A. hepatica are compiled.
Additional records of four species are reported, among them several new country
records of A. hepatica and the first record of A. gracilenta (Erichson,
1839) from Ukraine.
550:
Five
new species of Chinecallicerus from China (Coleoptera: Staphylinidae:
Aleocharinae: Geostibini).
Five
species of
Chinecallicerus
Assing, 2004 from China are
described and illustrated:
Chinecallicerus
formidabilis
nov.sp. (North Sichuan), C. glabriventris nov.sp. (West Yunnan), C.
reuteri nov.sp. (North Sichuan), C. feldmanni nov.sp. (North
Sichuan), and C. discrepans nov.sp. (North Sichuan). An updated key to
the Chinecallicerus species is provided. The distributions of the new
species and their respective closest relatives are mapped. Including the new
species, the genus now includes 18 species, nine of which have been recorded
from the Chinese province Yunnan, eight from Sichuan, and one from Qinghai. All
the species are known only from their respective type localities, fourteen of
them are represented exclusively by their respective holotypes.
551:
A
revision of the species of Blepharhymenus
of the Palaearctic and Oriental regions (Coleoptera: Staphylinidae:
Aleocharinae: Oxypodini).
Species
of Blepharhymenus Solier,
1849 had been reported from the Chilean Subregion (including the type species of
the genus), the Nearctic, Palaearctic, Ethiopian, and Australian regions,
suggesting that the overall distribution of this genus was remarkably
discontinuous. In order to clarify the status and generic assignment of the
Palaearctic species, types and additional material are revised, including also a
selection of species from Chile and North America. A study of this material
revealed that what had been regarded as Blepharhymenus in fact represents
various distinct lineages and that none of the species from the Palaearctic
Region belongs to the clade containing the type species of Blepharhymenus.
The following genus group taxa are newly described, revalidated, or elevated to
genus: Echidnoglossa Wollaston,
1864, revalidated (previously synonym of Blepharhymenus); Maurechidna
subgen.n. (subgenus of Echidnoglossa; type species: Echidnoglossa
ventricosa Quedenfeldt,
1881); Orechidna subgen.n. (subgenus of Echidnoglossa; type
species: Echidnoglossa hirthei sp.n.); Sinechidna subgen.n.
(subgenus of Echidnoglossa; type species: Blepharhymenus smetanai Pace,
2012); Syntomenus Bernhauer,
1939, stat.n. (previously subgenus of Blepharhymenus); Kortomenus
gen.n. (type species: Blepharhymenus koreanus Paśnik,
2001); Colusa Casey,
1885, revalidated (previously synonym of Blepharhymenus; type species Colusa
gracilis Casey,
1885). Echidnoglossa is discontinuously distributed across the
Palaearctic and northern Oriental regions and currently includes 19 species in
five subgenera. Syntomenus and the monotypical genus Kortomenus
are confined to the southern East Palaearctic and northern Oriental regions. All
the species from the Palaearctic and Oriental regions are (re-)described and
illustrated, six of them for the first time: Echidnoglossa (Echidnoglossa)
russa sp.n. (Egypt: Sinai Peninsula); Echidnoglossa (Orechidna)
hirthei sp.n. (Nepal); E. (O.) betzi sp.n.
(Thailand); E. (O.) artior sp.n. (Taiwan); E. (O.)
formosana sp.n. (Taiwan); Syntomenus laoticus sp.n. (Laos). Numerous
new combinations are proposed: the Palaearctic species previously attributed
to Blepharhymenus are all assigned to Echidnoglossa, Syntomenus,
or Kortomenus, and the binomen Ocalea dabensis (Pace,
2012), comb.n. (ex Blepharhymenus) is established.
The Nearctic species are tranferred to Colusa. The following synonymies
are proposed: Echidnoglossa corsica Mulsant
& Rey, 1875 = E.
sardoa (Scheerpeltz, 1954),
syn.n.; E. glabrata (Kiesenwetter,
1870) = E.
moczarskii (Scheerpeltz, 1954),
syn.n.; E. maghrebica (Fagel, 1960) =
E.
peyerimhoffi (Fagel, 1960),
syn.n.; E. meschniggi (Bernhauer,
1936) =
E.
elegans (Fagel, 1959),
syn.n. = E. guadalupensis (Fagel,
1959), syn.n.; Syntomenus
chinensis
(Bernhauer, 1939) = S. rougemonti
(Pace, 1999), syn.n. Lectotypes are
designated for Blepharhymenus moczarskii Scheerpeltz,
1954, Echidnoglossa paulinoi Skalitzky,
1884, and E. ventricosa Quedenfeldt,
1881. A checklist of the taxa distributed in the Palaearctic and Oriental
regions and a key to the species of Echidnoglossa is provided. The
distributions of Echidnoglossa species are mapped.
552:
Three
new species of Platyprosopus from Thailand, Cambodia, and Sierra Leone,
and additional records (Coleoptera, Staphylinidae, Staphylininae).
Three
species of Platyprosopus Mannerheim,
1830 are described and illustrated: P. latitarsalis nov.sp.
(Thailand); P. rossii nov.sp. (Cambodia); P. maximus nov.sp.
(Sierra Leone). Additional records of three species are reported primarily from
Cambodia, among them two new country records.
553:
A
revision of Geostiba of the West Palaearctic region. XXVII. New species
from Georgia and Kyrgyzstan, and additional records (Coleoptera, Staphylinidae,
Aleocharinae).
Five
species of Geostiba Thomson, 1858 from
Georgia and East Kyrgyzstan are described and illustrated: Geostiba (Tropogastrosipalia)
simplicicollis nov.sp. (Georgia: Kakheti); G. (Sibiota) kakhetiana
nov.sp. (Georgia: Kakheti); G. (S.) frischi nov.sp.
(Kyrgyzstan: Issyk-Kul); G. (S.) prominens nov.sp.
(Kyrgyzstan: Issyk-Kul); G. (S.) dentata nov.sp.
(Kyrgyzstan: Issyk-Kul). Based on a study of recently collected material, Geostiba
zerchei Pace, 1996 is moved
from the subgenus Sibiota Casey,
1906 to Sipalotricha Scheerpeltz,
1931; the male sexual characters of this previously doubtful taxon are described
and illustrated for the first time. Additional records of eight species are
reported from the Caucasus region.
554:
The
mother of synonyms: on the Meotica species of the Palaearctic Region
(Coleoptera, Staphylinidae, Aleocharinae, Oxypodini).
The
Holarctic genus Meotica Mulsant
& Rey, 1873 has been in a state of serious taxonomic confusion,
primarily because numerous species have been described without a study of the
sexual characters and because both intraspecific variation and distribution
ranges have been underestimated. Based on a revision of types and additional
material, the genus is represented in the Palaearctic region by 20 species, 18
of them distributed in the West Palaearctic and two in the East Palaearctic
regions. The identity of three additional species described from Northwest
Africa, whose type material is currently inaccessible, remains uncertain. One
genus-group and as many as 23 species-group synonymies are proposed for the
first time: Meotica Mulsant &
Rey, 1873 = Cryptusa Mulsant
& Rey, 1873, nov.syn.; Meotica filiformis (Motschulsky,
1860) = M. capitalis Mulsant
& Rey, 1873, nov.syn., = M. fageli Benick,
1953, nov.syn.; M. exilis (Gravenhorst,
1806) = M. simillima Benick,
1953, nov.syn.; M. pallens (Redtenbacher,
1849) = M. hanseni Benick,
1953, nov.syn.; M. filaria (Fauvel,
1898) = M. meridiogallica Scheerpeltz,
1954, nov.syn.; M. parasita Mulsant
& Rey, 1873 = M. winkleri Benick,
1953, nov.syn., = M. bosnica Scheerpeltz,
1954, nov.syn., = M. albanicola Scheerpeltz,
1969, nov.syn., = M.
aegyptia Pace, 1983, nov.syn.;
M. moczarskii Scheerpeltz,
1927 = M. finnmarchica Benick,
1953, nov.syn., = M.
romana Benick, 1953, nov.syn., =
M. albanica Benick, 1953, nov.syn.,
= M. marchica Benick,
1953, nov.syn., = M.
exigua Benick, 1953, nov.syn., =
M. alpina Benick, 1953, nov.syn.,
= M. testacea Benick,
1953, nov.syn., = M.
neapolitana Scheerpeltz, 1954,
nov.syn., = M. stockmanni Muona,
1978, nov.syn., = M.
anglica Benick, 1991, nov.syn.;
M. ochsi Benick, 1953
= M. punctulata Benick, 1953, nov.syn.;
M. normandi Benick, 1953 = M.
franzi Pace, 1983, nov.syn.; M.
caucasica Benick, 1953 = M.
decolor Assing, 2004, nov.syn.;
Apimela sabulicola (Bernhauer,
1914) = Meotica quadraticollis Scheerpeltz,
1954, nov.syn. Thus, Meotica now totals 58 synonyms outnumbering
the valid names by a factor of nearly three. The five most widespread species
alone account for 53 synonyms: M. moczarskii (19 synonyms), M.
filiformis (12), M. exilis (8), M. pallens (8), and M.
parasita (6). Meotica kuehnelti (Scheerpeltz,
1963) is transferred to the genus Tectusa Bernhauer,
1899. Since Tectusa kuehnelti (Scheerpeltz,
1963) is preoccupied by T. kuehnelti (Scheerpeltz, 1962), the name is replaced with Tectusa
killinica nov.nom. Lectotypes are designated for Meotica hoelzeli Benick,
1953, M. clavata Benick,
1953, M. simillima Benick,
1953, M. bosnica Scheerpetz,
1954, M. albanica Scheerpetz, 1967,
M. moczarskii Scheerpeltz,
1927, M. marchica Benick,
1953, M. alpina Benick,
1953, M. exigua Benick,
1953, M. roubali Benick,
1953, M. neapolitana Scheerpeltz,
1954, M. ochsi Benick, 1953,
M. smetanai Scheerpetz,
1967, M. normandi Benick,
1953, M. quadraticollis Scheerpeltz,
1954, and Atheta filaria Fauvel,
1898. A synonymic catalogue, as well as concise diagnoses and
illustrations of the sexual characters of all the species of which material was
available are provided.
555:
New
species and additional records of Ischnosoma from Georgia (Coleoptera,
Staphylinidae, Tachyporinae).
Four
species of Ischnosoma Stephens,
1829 from Georgia are described and illustrated, all of them belonging to the I.
spelaeum group: Ischnosoma acre nov.sp. (Southwest Georgia: Imereti
region), a close relative of I. solodovnikovi Schόlke,
2001 and I. molle Assing & Schόlke,
2017, I. convergens nov.sp. (Southwest Georgia: Adjara region: Meskheti
Range) and I. parallelum nov.sp. (Southwest Georgia: Adjara region:
Shavsheti Range), both closely allied to I. myops (Eppelsheim,
1880), and I. cuspidatum nov.sp. (Northeast Georgia: Kakheti region). For
comparison, new illustrations of I. solodovnikovi, I. molle, and I.
myops are provided. Additional records of four species are reported, among
them the first confirmed record of I thoracicum (Eppelsheim,
1880), a species with a most likely erroneous type locality. The distributions
of the new species and of I. solodovnikovi, I. molle, I. myops,
and I. thoracicum are mapped.
556:
Four
new species and additional records of Lomechusini from Mindanao, Philippines
(Coleoptera, Staphylinidae, Aleocharinae).
Drusilla
bucina nov.sp.,
D. holoserica nov.sp., Zyras (Zyras) rasilis nov.sp.,
and Z. (Z.) nigrorufus nov.sp. (all Philippines: Mindanao)
are described and illustrated. Six additional species, four of them unnamed, are
reported from Mindanao. Including the new species, 46 named species of
Lomechusini are currently known from the Philippines.
557:
On
the Staphylinidae of the Greek island Samothraki (Insecta, Coleoptera).
Aside
from individual records of two widespread species, the Staphylinidae of
Samothraki, a Greek island in the north of the Aegean Sea, had never been
studied. In April 2019, a field trip was conducted to Samothraki to investigate
the staphylinid fauna. A study of 1372 specimens collected during this field
trip yielded 123 species, 114 of them named and six of them endemic. Two species
and one subspecies are reported from Greece for the first time. Seven species
are described and illustrated: Apimela procera Assing
nov.sp. (Greece: Samothraki, Fthiotis; Turkey: Osmaniye) of the Aleocharinae;
Cyrtotyphlus samothracicus Assing
nov.sp. (endemic) and Metrotyphlus samothracicus Assing
nov.sp. (endemic) of the Leptotyphlinae; Cephennium samothracicum Assing
nov.sp. (endemic) and Scydmoraphes samothracicus Meybohm
nov.sp. (endemic) of the Scydmaeninae; Bryaxis samothracicus Brachat
nov.sp. (endemic) and Tychobythinus assingi Brachat
nov.sp. (endemic) of the Pselaphinae. A checklist of the species recorded from
Samothraki is provided. Samothraki is characterized by a relatively poor forest
fauna, but a high diversity of wetland species. It is compared to that of the
North Aegean Islands Chios, Lesbos, Samos, and Ikarνa regarding total diversity
and number of endemic species. In an appendix, Bryaxis pangeonicus Brachat
nov.sp. from Oros Pangιo (Northeast Greece) is described.
558:
A
new genus of Oxypodina from Kyrgyzstan (Coleoptera, Staphylinidae, Aleocharinae,
Oxypodini).
Typhlotectusa
nov.gen. frischi nov.sp. (East Kyrgyzstan: Issyk-Kul) of the subtribe
Oxypodina is described and illustrated. The new species was discovered in a
high-altitude habitat and is characterized by a slender habitus, its mouthparts,
strongly incrassate antennae, a short metatarsomere I, the shapes of the male
sternite VIII and the apical lobe of the paramere, as well as additionally by
the complete absence of eyes, reduced wings, and reduced pigmentation.
559:
The
first record of the subfamily Osoriinae from the Caucasus region (Coleoptera:
Staphylinidae).
Leptotyphlopsis
shavshetica
sp. nov. from the Shavsheti Range (Southwest Georgia: Adjara region), the first
representative of the subfamily Osoriinae and the first truly endogean species
of Staphylinidae to be recorded from the Caucasus region, is described and
illustrated. The distribution of this species is mapped.
560:
On
some Proteinus species of the Caucasus region, with description of a new
species from Georgia and a new synonymy (Coleoptera: Staphylinidae: Proteininae).
Proteinus
cavatus sp.
nov. (Southwest Georgia) of the P. longicornis group, a species with
conspicuous male secondary sexual characters, is described and illustrated.
Based on a study of type material, redescriptions and illustrations are provided
for Proteinus planicollis Reitter,
1905 and P. reflexicollis Reitter,
1905 and lectotypes are designated for both species. A new synonymy is proposed:
P. atomarius Erichson, 1840
= P. hiburnus Gistel, 1857,
syn. nov. The presence of two widespread species in Georgia is confirmed.
561:
A
new species of Parabolitobius from the Meskheti Range, Southwest Georgia,
with a new record of Bolitobius insignis (Coleoptera: Staphylinidae:
Tachyporinae).
Parabolitobius
meskheticus
sp. nov. (Southwest
Georgia: Meskheti Range), a micropterous and presumably endemic species, is
described, illustrated, and distinguished from allied and geographically close
spccies. Including the new species, Parabolitobius Li,
Zhao & Sakai, 2000 is currently represented in the Palaearctic region
by eleven species. A new record of the rare Bolitobius insignis is
reported from Southwest Georgia.
562:
On
the taxonomy and zoogeography of some Xantholinus species of the Caucasus
region, with descriptions of a new species from Georgia and two new synonymies
(Coleoptera: Staphylinidae: Staphylininae).
Xantholinus
(Calolinus)
mesheticus sp. nov. (Southwest Georgia: Adjara: Mesheti Range), a
micropterous species with a presumably restricted distribution, is described and
illustrated. Two synonymies are proposed: Xantholinus (Helicophallus)
variabilis Hochhuth, 1851 =
X. kirschenblati Bordoni, 1975,
syn. nov., = X. ceviki Anlaş,
2014, syn. nov. New records of five species of Xantholinus Dejean,
1821 are reported from Georgia. The distributions of six Caucasian species are
mapped.
563:
On
the Leptusa fauna of the Caucasus region II. A new species and additional
records from Georgia, and three new synonymies (Coleoptera: Staphylinidae:
Aleocharinae).
Leptusa
(Stictopisalia)
shavshetica sp. nov. (Southwest
Georgia: Adjara: Shavsheti Range), is described, illustrated, and distinguished
from similar species. Three synonymies are proposed: Leptusa (Roubaliusa)
trapezuntis Pace, 1983 = L.
giresunensis Sert, Turan & Kabalak,
2019, syn. nov.; Leptusa (Neopisalia) microphthalma Reitter,
1887 = L. circassica Bernhauer,
1935, syn. nov.; Leptusa (Stictopisalia) subcaucasica
Pace, 1983 = L. pseudocaucasica
Pace, 1983, syn. nov. Additional
records of six species of Leptusa Kraatz,
1858 are reported from Georgia. The distributions of three species are mapped.
The Caucasian Leptusa fauna currently includes 38 valid species in five
subgenera.
564:
A
revision of Othiini. XXI. Seven new species, a new synonymy, and additioonal
records, with a review of the Othius fauna of Georgia (Coleoptera:
Staphylinidae: Staphylininae).
Othius
derectus
sp. nov. (Southwest Georgia: Meskheti Range), O. unculatus sp. nov. (West
Georgia: Kutaisi env.), O. rasus sp. nov. (Southwest Georgia: Meskheti
Range), O. hetzeli sp. nov. (Taiwan),
O. brunkei sp. nov. (North
Vietnam), O. procerus sp. nov. (North Vietnam), O. devians sp.
nov. (North Vietnam), and O. bispinosus sp. nov. (China: Zhejiang) are
described, illustrated, and distinguished from allied species. One synonymy is
proposed: Othius lapidicola Mδrkel
& Kiesenwetter, 1848 = O. piceus Scriba,
1870, syn. nov. Additional records of twelve species are reported. The Othius
fauna of Georgia currently includes 16 described species. The distributions of
the Caucasian species recorded from Georgia and of the widespread, but
discontinuously distributed Othius lapidicola are mapped. Othius Stephens,
1829 now comprises a total of 137 species and subspecies.
565:
New
species and records of Lathrobium, with the first and southernmost record
of the genus from Vietnam (Coleoptera: Staphylinidae: Paederinae).
Six
species of the Holarctic genus Lathrobium
Gravenhorst,
1802 are described and illustrated: L.
austere sp. nov. (North Vietnam: Cao Băng Prov.); L. tsukubanum
sp. nov. (Japan: Honshu: Ibaraki Pref.); L. converrens sp. nov. (Japan:
Shikoku: Ehime Pref.); L. aperiens sp. nov. (Japan: Shikoku: Ehime
Pref.); L. toonicum sp. nov. (Japan: Shikoku: Ehime Prov.); L.
bigladiosum sp. nov. (Japan: Honshu: Fukui Pref.). Additional records of six
species are reported from China and Japan. The genus is now represented in the
Old World by as many as 620 species, 131 of which have been recorded from Japan.
An updated checklist of the Lathrobium species of Japan is provided.
566:
A
revision of Palaearctic Lobrathium. VII. A new species from Greece, a
revalidation, and additional records (Coleoptera: Staphylinidae: Paederinae).
Lobrathium
moreanum
sp. nov. (Greece: Pelopσnnisos: Menalo Oros), a close relative of L.
multipunctum
(Gravenhorst,
1802), is described and illustrated. Lobrathium
gallienii (Fagniez,
1917), previously a synonym of L.
multipunctum,
is revalidated. Additional records of seven species from Turkey, Iran, and China
are reported. The genus is now represented in the Palaearctic region by 153
species.
568:
Monograph
of the Staphylinidae of Crete (Greece). Part I. Diversity and endemism
(Insecta:Coleoptera).
With
a geological history of long isolation, fragmentation, partial submersion, and
subsequent rising, characterized by a remarkable diversity in topology, geology,
and habitats, and the fifth-largest island of the Mediterranean, Crete has
provided an ideal setting for speciation and developing a unique and diverse
Staphylinidae fauna. Although beginning already in the middle of the 19th
century, the exploration of this fauna gained momentum only recently. Based on a
critical revision of literature data, on material collected during several
recent field trips, and on additional previously unpublished records, a
checklist of the Staphylinidae of Crete is compiled, including as many as 397
named species, with 115 of them reported from the island for the first time and
with 48 new species described in the second part of the monograph. Additional
unidentified and unnamed species, as well as species doubtfully or erroneously
recorded from Crete are listed separately. The systematic, zoogeographic, and
ecological composition of the Staphylinidae fauna is characterized with a
special focus on the endemics. The latter are represented by 111 named and at
least ten unnamed species, account for nearly one-third of the fauna, and mainly
belong to the subfamilies Aleocharinae, Scydmaeninae, Pselaphinae, Paederinae,
Leptotyphlinae, Staphylininae, and Omaliinae. More than half of them are even
locally endemic. The genera with the greatest number of endemic species are Cephennium
Mόller & Kunze, 1822 of the
Scydmaeninae (twelve named species) and Geostiba Thomson,
1858 of the Aleocharinae (nine named species). More than two-thirds of the
endemic species are epigeic inhabitants of forests, bush and shrub habitats,
grassland, and wetlands, approximately one-fourth is confined to deeper soil
strata or endogean, and 8 % are myrmephilous. Slightly more than half (59
species; 53 %) of the endemics are most likely the result of in-situ radiation
(18 lineages), the remainder of vicariance with mainland species. A comparison
with other Mediterranean islands revealed that (a) total diversity of the Cretan
fauna is significantly greater than that of other East Mediterranean islands
except Corfu, but much lower than that of Corsica, Sardinia, and Sicily, (b)
regarding the number of endemic species Crete ranks third behind Corsica and
Sardinia, and (c) the rate of endemism is greater in Crete than in any other
Mediterranean island. Explanations for the observed differences in the diversity
and composition of the faunas of the islands are discussed. Stichoglossa
graeca Bernhauer,
1905,
a species of which previously only the lectotype was known, is redescribed and
illustrated. Six synonymies are proposed and one name is revalidated: Phloeocharis
longipennis Fauvel,
1875 = P. hummleri Bernhauer,
1915, syn. nov.; Phytosus balticus Kraatz,
1859
= P. holtzi Bernhauer,
1935,
syn. nov.; Atheta nigra (Kraatz,
1856) = A. biroi Scheerpeltz,
1964; syn. nov.; Domene stilicina (Erichson,
1840) = D. lohseiana Bordoni,
1977, syn. nov.; Oedichirus rubronotatus Pic,
1903, revalidated = O. reitteri Bernhauer,
1908, syn. nov.; Pseudolathra quadricollis (Fauvel,
1875)
= P. cretensis Bordoni,
1986,
syn. nov. A lectotype is designated for Stichoglossa
graeca Bernhauer,
1905. Two
species are reported from Greece for the first time.
569:
Monograph
of the Staphylinidae of Crete (Greece). Part II. Descriptions of new species
(Insecta:Coleoptera).
One
genus, Cretotyphlus Assing
gen. nov., and 48 species of Staphylinidae from the Greek island Crete, 47 of
them island endemics or even locally endemic, are described and illustrated: Amischa
cretica Assing spec. nov., Atheta
(Anopleta) digitalis Assing
spec. nov., Hydrosmecta insularum Assing
spec. nov. (Crete, Ikarνa, Lesbos, Samothraki), Geostiba (Sipalotricha)
inexsecta Assing spec. nov.
(region to the east-southeast of Rethimno), Cousya candica Assing
spec. nov. (East Crete: Dikti), Oxypoda (Bessopora) bimontium
Assing spec. nov. (Dikti,
Psiloritis), O. (Mycetodrepa) retunsa Assing
spec. nov., and Typhlocyptus creticus Assing
spec. nov. of the Aleocharinae; Geomitopsis cretica Assing
spec. nov. (East Crete) of the Osoriinae; Allotyphlus (Moreotyphlus)
foedatus Assing spec. nov.
(West Crete), Kenotyphlus virgatus Assing
spec. nov. (West Crete), Cretotyphlus hamatus Assing spec. nov. (East Crete: Dikti), C. chanianus Assing
spec. nov. (region to the southwest of Chania), and C. idanus Assing
spec. nov. (Central Crete: Psiloritis) of the Leptotyphlinae; Pseudobium
creticum Assing spec. nov. of
the Paederinae; Gabrius candicus Assing
spec. nov. of the Staphylininae; Cephennium arcuatum Assing
spec. nov. (East Crete), C. thripticum Assing
spec. nov. (East Crete: Thripti), C. selinonum Assing
spec. nov. (Southwest Crete), C. meybohmi Assing spec. nov. (West Crete), C. idanum Assing
spec. nov. (Central Crete: Psiloritis), C. sinuosum Assing
spec. nov. (East Crete), C. fortespinosum Assing spec. nov. (East Crete), C. hamulatum Assing
spec. nov. (East Crete: Dikti), C. curvatum Assing
spec. nov. (East Crete), C. selenanum Assing
spec. nov. (East Crete: Selena Oros), C. latius Assing spec. nov. (Central Crete: Psiloritis), C.
chanianum Assing spec. nov.
(West Crete), Euconnus (Tetramelus) zakrius Meybohm
spec. nov. (East Crete), Stenichnus (Stenichnus) brachati Meybohm
spec. nov., S. (S.) orientalis Meybohm spec. nov. (East Crete), S. (S.) aegialioides
Meybohm spec. nov., Leptomastax
cretica Meybohm spec. nov., and
L. thriptica Meybohm spec.
nov. (East Crete: Thripti) of the Scydmaeninae; Afropselaphus doernfeldae
Brachat spec. nov. (West Crete:
Lefka Ori), A. assingi Brachat
spec. nov. (West Crete: Lefka Ori), A. diktianus Brachat spec. nov. (East Crete: Dikti), A. thripticus Brachat
spec. nov. (East Crete: Thripti), Amauronyx chanianus Brachat
spec. nov. (West Crete), A. occidens Brachat
spec. nov. (West Crete), A. askifouicus Brachat
spec. nov. (West Crete), Bryaxis meybohmianus Brachat spec. nov. (West Crete), Bythinus creticus Brachat
spec. nov. (Central
Crete: Psiloritis), Euplectus assingi Brachat
spec. nov. (Crete),
Faronus meybohmi Brachat
spec. nov. (West
Crete: Lefka Ori), F. lefkamontium Brachat
spec. nov. (West
Crete: Lefka Ori), F. albimontium Brachat
spec. nov. (West Crete: Lefka Ori), and Tychus chanianus Brachat
spec. nov. (West Crete) of the Pselaphinae; Sepedophilus creticus Schόlke
spec. nov. of the Tachyporinae. Keys to the species of Cephennium Mόller
& Kunze, 1822 and Stenichnus Thomson,
1859 of Crete are provided. A lectotype is designated for Amauronyx
paganettii Blattnύ & Blattnύ,
1916.
570:
On
the taxonomy and zoogeography of some West Palaearctic Cypha species
(Coleoptera: Staphylinidae: Aleocharinae).
Three
species are (re-)described and illustrated: Cypha squamipennis (Fauvel,
1902), C. carinthiaca (Scheerpeltz,
1958), and C. bisinuata sp. nov. (South Italy). Based on an
examination of types and additional material, C. hanseni (Palm,
1949) is removed from synonymy with C. seminulum (Erichson,
1839) and synonymized with C. pulicaria (Erichson,
1839). A revision of material of species of the newly established C.
spathulata group (including C. spathulata Assing,
2007, C. graeca Assing,
2004, C. lindbergi (Palm,
1935), and C. bisinuata) revealed an allopatric distribution pattern in
the Mediterranean and Caucasus regions. The distributions of these species are
mapped. The male primary sexual characters of C. seminulum, C.
pulicaria, and of the species of the C. spathulata group are
illustrated. Cypha seminulum is reported from Portugal and Georgia for
the first time.
571:
Revision
of the Apimela species of the Palaearctic Region with a focus on the West
Palaearctic fauna, and with additional notes on the genus Franzidota
(Coleoptera: Staphylinidae: Aleocharinae: Oxypodini).
Based
on a revision of types and additional material, the oxypodine genus Apimela
Mulsant & Rey, 1874 is
represented in the Palaearctic region by 29 described species, seven of them
confined to the West Palaearctic including Middle Asia, 21 to the East
Palaearctic, and one widespread in the southern East Palaearctic and Oriental
regions. Two additional species of which neither type material nor illustrations
were available remain of uncertain status. Most of the species are
(re-)described and/or illustrated, nine of them for the first time: Apimela
mutata sp. nov. (=
A. macella auctt.) (widespread
in the West Palaearctic region); A. wunderlei sp. nov. (South Italy); A.
graeca sp. nov. (Greece); A. kirghisica sp. nov. (Kyrgyzstan); A.
ussurica sp. nov. (Russian Far East); A. baculata sp. nov. (China:
Yunnan); A. bilobata sp. nov. (China: Zhejiang, Sichuan, Yunnan); A.
auriculata sp. nov. (China: Yunnan); A. lamellata sp. nov. (North
Vietnam). Two genus-group and four species-group synonymies are proposed: Apimela
Mulsant & Rey, 1874 = Parapimela Cameron,
1939, syn. nov., = Himalkompsusa Pace,
2006, syn. nov.; Apimela macella (Erichson,
1839) = A. pallens (Mulsant &
Rey, 1852), syn. nov., = A. mulsanti (Ganglbauer,
1895), syn. nov.; A. indica (Cameron,
1939) = A. assamensis (Cameron,
1939), syn. nov.; A.
chinensis Pace, 1999 = A.
tibetana Pace, 2012, syn. nov.
The genus-group synonymies result in the following binomina: Apimela
indica (Cameron, 1939), comb.
nov.; A. persimilis (Cameron, 1939), comb. nov.; A. nepalicola (Pace, 2006), comb.
nov.; A. sinica (Pace,
2012), comb. nov.; A. hartmanni (Pace, 2006),
comb. nov.; A. taiwanensis (Pace, 2010), comb. nov.; A.
morvani (Pace,
1992), comb. nov. Apimela
lucidula Pace,
1992 belongs
to an oxypodine genus of unknown identity and is excluded from Apimela.
For various reasons (type material not accessible or not found; descriptions
based on females only), several species remain of doubtful identity and partly
uncertain generic assignment. Apimela is represented in the Palaearctic
region by two distinct clades distinguished primarily by the male and female
sexual characters, the A. macella lineage (seven species) and the A.
mutata lineage (22 species), both of them distributed across the Palaearctic
region. Most Apimela species inhabit the banks of rivers and streams with
substrates composed of coarse sand and fine gravel in mountainous regions. An
updated catalogue of the Apimela species of the world and a key to the
species of the West Palaearctic region including Middle Asia are provided. The
distributions of ten species are mapped. In an appendix, the genus Franzidota
Pace, 1982 is moved from Oxypodina to Meoticina, and F.
formosana sp. nov. (Taiwan) and F. gilva sp. nov. (Taiwan) are described and illustrated.
572:
On
the taxonomy and zoogeography of Paederus. V. Two new species from Laos
and China, a new synonymy, new subgeneric assignments, and new records from the
Palaearctic region (Coleoptera: Staphylinidae: Paederinae).
Paederus
laoticus
sp. nov. (Northwest Laos: Bokeo province) and P. (Harpopaederus)
willersi sp. nov. (China: Sichuan) are described and illustrated. A new
synonymy is proposed: Paederus riparius (Linnaeus,
1758) = P. melanurus Aragon,
1830, syn. nov. Paederus kambaitiensisis Scheerpeltz,
1965 is assigned to the subgenus Heteropaederus Scheerpeltz, 1957 and P. tamulus Erichson,
1840, previously in Eopaederus Scheerpeltz,
1957, is removed from this subgenus and treated as Paederus incertae
sedis. The aedeagus of P. kambaitiensis is illustrated for the first
time. New records of nine species are reported, among them a new record from the
Palaearctic region, two from China, and two new province records from China. The
revised distributions of eight species of the subgenus Harpopaederus Scheerpeltz, 1957 are mapped.
573:
On
the Tetrasticta fauna of Laos (Coleoptera: Staphylinidae: Aleocharinae:
Aleocharini).
Three
species of Tetrasticta Kraatz,
1857 are described and illustrated, all of them from the same locality in Bokeo
province, Northwest Laos: Tetrasticta varia spec. nov.; T. crassa
spec. nov.; T. puncticeps spec. nov. The habitus and primary sexual
characters of T. brevipennis (Bernhauer,
1903) are figured. Aside from the new species, four species are reported from
Laos, three of them for the first time. Thus, the currently known Tetrasticta
fauna of Laos is composed of seven species. Tetrasticta brevipennis is
reported from Malaysia, the Indian province Uttarakhand, and the Chinese
province Guangdong, and T. bobbii Zheng
& Zhao, 2014 from Myanmar for the first time. In an addendum, Tetrasticta
borneana spec. nov. (Malaysia: Sabah) is described and illustrated. The
genus now includes a total of 20 species, except for one Aftrotropical species
all of them distributed in the southern East Palaearctic and Oriental regions.
574:
A
revision of Palaearctic and Oriental Rugilus VII.Two new species from
China and Laos, and additional records (Coleoptera: Staphylinidae: Paederinae).
Two
species of Rugilus Leach,
1819 are described and illustrated: Rugilus (Rugilus) biapicalis spec. nov. (China: Yunnan); Rugilus
(Eurystilicus)
laoticus spec. nov. (North Laos: Bokeo province). Additional records of five
species of the subgenus Eurystilicus Fagel, 1953 are reported, among them two first records from
Vietnam. The genus now includes a total of 255 species worldwide, 104 of which
are distributed in the Palaearctic and Oriental regions. As many as 36 species,
most of them endemic, have been reported from China.
575:
Three
new myrmecophiles of the tribes Sahlbergiini and Aenictoteratini from Laos and
Malaysia (Borneo) (Coleoptera: Staphylinidae: Aleocharinae).
Malayloeblius
laoticus
spec. nov. (Northwest Laos) and M. tensus spec. nov. (Borneo: Sabah) of
the Sahlbergiini, and Dentaphila iniqua spec. nov. of the Aenictoteratini
(Northwest Laos) are described, illustrated, and distinguished from previously
described species of the respective genera. Two additional records of Malayloeblius
borneensis Pace, 2014 are
reported from Borneo. The host ants of the new species are unknown, but
available evidence suggests that these species may be associated with army ants.
576:
On
the taxonomy of Anomognathus Solier,
1849: the first records of the genus from Laos and a new synonymy (Coleoptera:
Staphylinidae: Aleocharinae).
Anomognathus
laoticus
sp. nov. (Northwest Laos: Bokeo province) and A. triangularis sp. nov.
(Northeast Laos: Houa Phan province), the first representatives of the genus to
be recorded from Laos, are described and illustrated. A new synonymy is
proposed: Placusa nitida Fauvel, 1900 = Anomognathus ispartaensis
Φrgel, Avci & Φzek, 2019; Placusa nitida is reported from Spain,
Turkey, and Israel for the first time. The genus Anomognathus Solier,
1849 currently includes 27 valid species distributed in all major zoogeographic
regions, except for the Afrotropical region.
577:
On
the taxonomy and zoogeography of Anaulacaspis Ganglbauer,
1895 and Borboropora Kraatz,
1862: new species, new records, and the first records of both genera from Laos
(Coleoptera: Staphylinidae: Aleocharinae: Falagriini).
Anaulacaspis
spinosa
sp. nov. (Laos:
Bokeo province), A. aberrans sp. nov. (Laos: Bokeo and Xieng Khouang
provinces), and Borboropora laotica sp. nov. (Laos: Bokeo province) are
described and illustrated. Additional records of nine previously described
species of Anaulacaspis Ganglbauer,
1895
and one of Borboropora Kraatz,
1862 are reported, among them four new country records from the West Palaearctic
region and one first record from Laos and the Oriental region. Anaulacaspis
is recorded from Laos, Borboropora from Laos and the Oriental region for
the first time.
578:
On
the taxonomy and zoogeography of Chinecallicerus VIII. New species and
records, with the first record of the genus from Laos and from outside China
(Coleoptera: Staphylinidae: Aleocharinae: Geostibini).
Four
species of Chinecallicerus Assing, 2004 are described and illustrated: Chinecallicerus
dispersus sp. nov. (China: West Sichuan, West Yunnan); C. unituberculatus
sp. nov. (China: Sichuan: Gongga Shan); C. dabaicus sp. nov. (China:
Hubei: Daba Shan); C. laoticus sp. nov. (Northwest Laos: Bokeo province).
The previously unknown male sexual characters of C. formidabilis Assing,
2019 and the spermathecae of C. serratus Assing, 2009, C. pinnatus
Assing, 2017, and C. laevigatus Assing, 2006 are illustrated for the
first time. These species, all of them previously known only from their
respective holotypes, are reported from additional localities in Sichuan and
Yunnan. Chinecallicerus laoticus represents the first record of the genus
from Laos, from outside China, and from the whole of the Oriental region, C.
dabaicus the first record from the Chinese province Hubei and the
easternmost record of the genus. Enormous intraspecific variation was observed
for C. dispersus. Including the new species, the genus now includes 22
species, ten of which have been recorded from the Chinese province Yunnan, ten
from Sichuan, one from Qinghai, one from Hubei, and one from Laos. As many as 17
species are known only from their respective type localities, twelve of them are
represented exclusively by their respective holotypes. A checklist and a newly
designed key to species are provided. The distributions of the genus as a whole
and of seven individual species are mapped.
579:
Revision
of Amarochara VI. New species and records from the Palaearctic and
Oriental Regions (Coleoptera: Staphylinidae: Aleocharinae: Aleocharini).
Three
species of Amarochara Thomson, 1858
are described and illustrated: Amarochara sinuosa sp. nov. (North Laos,
North Vietnam); A. bifurcata sp. nov. (Northeast Laos: Houa Phan
province); A. spinosula sp. nov. (Northeast Laos: Houa Phan province).
Additional records of five previously described species are reported, among them
new country and new province records. The genus now includes a total of 50
species.
580:
Revision
of the Alevonota species of the Palaearctic region IV. Three new species
from Sichuan, China (Coleoptera: Staphylinidae: Aleocharinae: Geostibini).
Three
species of Alevonota Thomson,
1858, all of them from West Sichuan, China, are described and illustrated: Alevonota
pliciventris sp. nov. (Gongga Shan); A. ambulans sp. nov. (Shalui
Shan); A. tenuilobata (Erlang Shan). The genus is now represented in the
Palaearctic region by 49 species, 28 of them West Palaearctic and 21 East
Palaearctic.
581:
On
Orsunius IV. Three new species, new records, and the first record of the
genus from the Australian region (Coleoptera: Staphylinidae: Paederinae:
Medonina).
Three
species of the genus Orsunius Assing,
2011 are described and illustrated: Orsunius tortus sp. nov.
(Thailand) and O. frischi sp. nov. (Indonesia: Jawa Barat), both of them
belonging to the O. wunderlei group, as well as O. maindai sp.
nov. (Indonesia: Papua), the first record of the genus from the Australian
region. Additional records of nine previously described species are reported,
among them five new country records from Thailand (2), China (1), Hong Kong (1),
and Cambodia (1). The genus now includes 24 named extant species distributed in
the southern East Palaearctic, the Oriental, and the northern Australian
regions.
582:
A
new species of Lathrobium from Shikoku, Japan (Coleoptera: Staphylinidae:
Paederinae).
Lathrobium
sugitatense
sp. nov. (Japan: Shikoku: Ehime: Mt. Sugitate) is described, illustrated, and
compared with its close relative, L. kamezawai Watanabe, 2005. New
figures are provided for L. kamezawai. Based on several conspicuous
synapomorphies, L. sugitatense and L. kamezawai are hypothesized
to represent adelphotaxa. The Lathrobium fauna of Japan is now composed
of 134 species, 24 of which are distributed in Shikoku.
583:
On
the Cephennium
fauna
of Greece
(Coleoptera,
Staphylinidae, Scydmaeninae).
Thirteen
species of Cephennium Mόller & Kunze, 1822 from Greece are described and
illustrated: C. dilatatum nov.sp. (Kavala: Oros Pangιo), C. olympicum
nov.sp. (Pieria: Olympos), C. pellucidum nov.sp. (Fthiotis/Evritania:
Oros Timfristos), and C. brachati nov.sp. (Flσrina) of the C.
perispinctum group; C. aciferum nov.sp. (Pelopσnnisos, Thessalia,
Etolia-Akarnania, Kefalonia, Corfu), C. sagittatum nov.sp. (Ipiros), and C.
linguatum nov.sp. (Levkas) of the C. jonicum group; C. cornutum
nov.sp. (Fthiothis: Oros Timphristos) of the C. corruptum group; C.
vitsiense nov.sp. (Flσrina: Oros Vitsi); C. vellicans nov.sp. (Ioαnnina:
Oros Gramos); C. catax nov.sp. (Thessalia: Oros Ossa); C. dramanum
nov.sp. (Drama); C. fibuliferum nov.sp. (Fthiothis, Pelopσnnisos). Cephennium
jonicum
Holdhaus, 1908 and C. petraeum
Apfelbeck, 1911, previously subspecies of C.
jonicum, are treated as distinct species. Based on drawings by the late
Claude Besuchet, the aedeagi of several Cephennium species described
prior to 1910, of which published illustrations had not been available, are
figured. New records of two previously described species are reported. Including
the newly described taxa, the Cephennium fauna of Greece currently
includes 35 named species, twelve from Crete (endemic), one from Karpathos
(endemic), one from Samothraki (endemic), one from Levkas, and the remainder
from the mainland, the Pelopσnnisos, and the Ionian Islands Corfu, Levkas, and
Kefalonia. A catalogue of the Cephennium species recorded from Greece is
provided.
584:
On
the Cephennium
fauna
of the Iberian Peninsula and the Atlantic Islands
(Coleoptera,
Staphylinidae, Scydmaeninae).
Based
on a study of Iberian and Madeiran material of Cephennium Mόller
& Kunze, 1822, as well as on unpublished notes compiled by the late
Claude Besuchet, 38 named species are recognized in Iberian Peninsula to the
south of the Spanish border with France and Andorra, and three in Madeira, the
only Atlantic archipelago from where the genus has been recorded. The species
are attributed to eight species groups. With three exceptions (male unknown,
type material lost, or identity doubtful), illustrations or references to
recently published figures are provided for all the species of the Iberian
Peninsula and Madeira. Seventeen species are described for the first time: C.
constrictum nov.sp. (Spain: Cataluρa: Sierra del Cadi) of the C.
perispinctum group; C. validum nov.sp. (Northwest Spain, North
Portugal), C. conlectior nov.sp. (North Portugal: Serra da Estrela), C.
algarvense nov.sp. (Portugal), and C. lompei nov.sp. (Madeira proper)
of the C. validum group; C. filabresicum nov.sp. (Spain: Andalucνa:
Sierra de los Filabres) and C. espunae nov.sp. (Spain: Murcia: Sierra de
Espuρa) of the C. caecum group; C. tensum nov.sp. (Spain:
Alicante: Sierra de Aitana) and C. andujari nov.sp. (Spain: Alicante:
Sierra de Aitana) of the C. tensum group; C. tenuissimum nov.sp.
(Spain: Murcia: Sierra de Espuρa) of the C. tenuissimum group; C.
alacantae nov.sp. (Spain: Alicante), C. cujonicum nov.sp. (Spain:
Castilla-La Mancha: Sierra de Segura), C. verrens nov.sp. (Spain: Andalucνa:
Sierra de Segura), C. segurae nov.sp. (Spain: Castilla-La Mancha: Sierra
de Segura), C. nevadae nov.sp. (Spain: Andalucνa: Sierra Nevada), C.
cazorlae nov.sp. (Spain: Andalucνa: Sierra de Cazorla), and C. malagae nov.sp.
(Spain: Andalucνa: Malaga) of the subgenus Macroderus Croissandeau,
1894. Two subspecies of C. pygmaeum Saulcy,
1870
are regarded as doubtful. One species is recorded from Spain for the first time.
A catalogue of the named Cephennium species recorded from the Iberian
Peninsula and Madeira is provided. The distributions of 23 species are mapped.
Including the new taxa, the genus is now represented by 251 named species.
585:
On
the taxonomy of Parocyusa, Tectusa, and miscellaneous genera of
Oxypodina
(Coleoptera:
Staphylinidae: Aleocharinae: Oxypodini).
A
comparative morphology-based study of types and additional material of species
previously assigned to Tetralaucopora
Bernhauer,
1928,
Tectusa Bernhauer,
1899, and
various other genera of Oxypodina results in numerous taxonomic changes. Parocyusa
Bernhauer,
1902,
previously a synonym of Tectusa,
is
revalidated. The genus has a Holarctic distribution and currently includes 33
described species, 20 species in the West Palaearctic (including Middle Asia),
twelve in the East Palaearctic, and two species (one of them probably native in
the West Palaearctic) in the Nearctic region. The validity of two species, one
from North Europe and one from Japan, requires confirmation and one name is
treated as a tentative synonym. Ten of the West Palaearctic species are
micropterous or submacropterous and have restricted distributions, whereas the
remainder is macropterous and more or less widespread. Tectusa presently
includes 41 described species, all of them micropterous and with restricted
distributions, with the vast majority confined to mountain ranges in the
Balkans. The genus is most likely polyphyletic. However, with the type material
of several species (including the type species of available genus-group names)
currently inaccessible, this issue will have to be addressed in a future
revision. New combinations are proposed for all the species previously in Tetralaucopora
and some previously in Tectusa or other genera, nine names are
synonymized, and one name is preserved according to Article 23.9 of the Code
(ICZN 1999): Parocyusa
Bernhauer,
1902 = Tetralaucopora
Bernhauer,
1928, resyn.;
Parocyusa antennata
(Eppelsheim,
1878), comb.n.; P. baicalensis (Eppelsheim,
1893), comb.n. = beijingensis (Pace,
1999), syn.n., = hebeiensis (Pace,
1999), syn.n.; P. bicolorata
(Assing,
2007), comb.n.; P. caligula (Assing,
1996), comb.n.; P. carnica Lohse,
1988; P. championi (Cameron,
1939), comb.n.; P. crebrepunctata (Strand,
1962), comb.n.; P. fuliginosa (Casey,
1906), comb.n.; P. germana (Cameron,
1939), comb.n. = rougemonti (Pace,
1986), syn.n.; P. hartmanni
(Pace,
2013), comb.n. (ex Ocalea); P. holdhausi
(Bernhauer,
1902); P. japonica
(Cameron,
1933), comb.n.; P. knabli
(Bernhauer,
1914) = tirolensis
Scheerpeltz,
1958, syn.n.;
P.
lebedevi
(Bernhauer,
1928), comb.n. = bucharica (Bernhauer,
1928), syn.n.; P.
longicollis
(Eppelsheim,
1889), comb.n.; P. longitarsis (Erichson,
1839), comb.n., nomen protectum = attenuata (Stephens,
1832), nomen oblitum, =
syriaca (Saulcy,
1865), syn.n.; P. montana (Kraatz,
1856), comb.n.; P. ripicola (Cameron,
1939), comb.n.; P. rubicunda
(Erichson,
1837), comb.n. = cingulata (Kraatz,
1856), syn.n.; P. schuelkei (Assing,
1996), comb.n.; P. strupiiana
Scheerpeltz,
1958; P. subcyanea
(Cameron,
1939), comb.n.; P.
yakouensis
(Pace,
2010), comb.n. (ex Ocyusa); P. yunnanensis (Pace,
1993), comb.n.; Ocyusa
fuscobrunnea
(Cameron,
1939), comb.n.; Cousya
quadrisulcata (Bernhauer,
1935) = nepalensis Pace,
2006, syn.n.; Oxypoda sinonigra
(Pace,
2017), comb.n.;
Oxypoda besucheti Focarile,
1982. Numerous species of Parocyusa and Tectusa are
(re-)described and illustrated, among them 19 new taxa (nine of Parocyusa
and ten of Tectusa): Parocyusa dilatata
sp.n. (China: Yunnan); P. gilvipennis sp.n. (Kazakhstan); P. gonggaica
sp.n. (China: Sichuan); P. gracillima sp.n. (South Turkey); P. kahleni
sp.n. (South Slovenia); P. maculipennis
sp.n. (Northwest Iran); P. matarata
sp.n. (South Iran); P. spinosa
sp.n. (China: Gansu); P. virilis
sp.n. (Caucasus region: Russia, Armenia); Tectusa acrilobata
sp.n
(Erimanthos); T. apanoica
sp.n.
(Menalo); T. chelmosica sp.n. (Aroania); T. fimbriata sp.n.
(Panahaiko); T. latilobata sp.n. (Giona, Iti, Parnassos); T. menaloica
sp.n.
(Menalo); T. parnonica
sp.n.
(Parnon); T. pauli
sp.n.
(Aroania); T. taygetana
sp.n.
(Taygetos); T. vodiasa
sp.n.
(Panahaiko). All the newly described Tectusa species are distributed in
Greece and, except for T. latilobata, all of them in the Pelopσnnisos.
The Carpathian and one Anatolian species previously attributed to Tectusa
are moved to the genus Duplocyusa gen.n. (type species Oxypoda
uludaghensis Fagel,
1971),
with the Carpathian species in the subgenus Carpocyusa subgen.n. (type
species Tectusa
nigromontis
Zerche,
2007):
Duplocyusa (Duplocyusa) uludaghensis
Fagel,
1971,
comb.n.; Duplocyusa (Carpocyusa) nigromontis
(Zerche,
2007), comb.n.; D. (C.) bucegiensis
(Zerche,
2007), comb.n.;
D.
(C.) transsylvanica (Zerche,
2007), comb.n.;
D.
(C.) rosenauensis (Zerche,
2007), comb.n.;
D.
(C.) ceahlauensis (Zerche,
2007), comb.n.;
D.
(C.) rodnaensis (Zerche,
2007), comb.n.;
D.
(C.) calimaniensis (Zerche,
2008),
comb.n.;
D.
(C.) iucasensis (Zerche,
2008),
comb.n.
In an appendix on miscellaneous genera of Oxypodina, the following revalidation,
synonymies, and new combinations are proposed, and five species are described: Eurylophus
J.
Sahlberg, 1876,
revalidated = Drepasiagonusa Pace,
2012, syn.n.; Eurylophus grandiceps
J.
Sahlberg, 1876, comb.n. = feldmanni
(Assing,
2018), syn.n.; Eurylophus smetanai (Pace,
2012), comb.n.; E. procerus (Assing,
2018), comb.n.; E. tibeticus (Assing,
2018), comb.n.; E. angulatus (Assing,
2018), comb.n. (all Eurylophus species except E. grandiceps
ex Drepasiagonusa); Cousya
lobifera Pace,
2015 (probably belonging to an undescribed genus); Cousya
luteipennis sp.n. (Uzbekistan); Parocalea
tenebricosa sp.n. (Russian Far East); Trichoglossina
taiwafimbriata (Pace,
2010), comb.n. (ex Oxypoda); Trichoglossina
retunsa sp.n. (China: Yunnan); T.
tricuspidata sp.n. (China: Yunnan); Chinecousya
globosa sp.n. (China: Yunnan). Lectotypes are designated for Chilopora antennata Eppelsheim,
1878, C. baicalensis Eppelsheim,
1893, C. ripicola Cameron,
1939, and
C. championi Cameron,
1939. The distributions of Parocyusa rubicunda, a species with evidently
parthenogenetic populations in large parts of its range, and of 17 species of Tectusa
are mapped. Numerous new records are reported.
586:
On
the Lomechusini of the Palaearctic and Oriental regions. XXVIII. New genera,
species, and records, with a focus on the fauna of Laos and Borneo (Coleoptera:
Staphylinidae: Aleocharinae).
Based
on a morphological study of material of 14 genera of Lomechusini from the
Oriental and East Palaearctic regions, primarily from Laos and Borneo, and of
some specimens from Irian Jaya, numerous species are described and illustrated,
78 of them for the first time: Orphnebius abbreviatus sp. nov. (Laos), O.
crassisetosus sp. nov. (Laos), O. taleatus sp. nov. (Borneo), O.
geminus sp. nov. (Borneo), O. sextuberculatus sp. nov. (Borneo), O.
retornatus sp. nov. (Borneo), O. derectus sp. nov. (Borneo), O.
comatus sp. nov. (Laos), O. calvus sp. nov. (Laos), O. baculifer sp.
nov. (Borneo), O. unguicus sp. nov. (Borneo), O. curvatus sp. nov.
(Borneo),
O. varicornutus sp. nov. (Laos),
O. biacer sp. nov. (Borneo),
O. conifer sp. nov. (Borneo), O. mollis sp. nov. (Borneo), O.
linguis sp. nov. (Borneo), O. sinulinguis sp. nov. (Borneo), O.
cuspilinguis sp. nov., O. colitus sp. nov. (Borneo),
O. tensus sp. nov. (Borneo), and O. pressatus sp. nov. (Borneo) of
the Orphnebius hauseri group; Orphnebius (Deroleptus)
transfigens sp. nov. (Borneo), O. (D.) aciformis sp.
nov. (Borneo), O. (D.) acifer sp. nov. (Borneo), O. (D.)
quadricultratus sp. nov. (Borneo), O. (D.) floreni sp.
nov. (Borneo), O. (D.) obelifer sp. nov. (Borneo), O. (D.)
nodatus sp. nov. (Borneo), O. (D.) filiformis sp. nov.
(Borneo), O. (D.) plexus sp. nov. (Borneo), O. (D.)
tuberculifer sp. nov. (Borneo), O. (D.) tuberosus sp.
nov. (Borneo), O. (D.) granulosus sp. nov. (Borneo), O. (D.)
verrucosus sp. nov. (Borneo), O. (D.) botulus sp. nov.
(Borneo), O. (D.) pinnatus sp. nov. (Borneo),
O. (D.) bicarinatus sp. nov. (Borneo),
O. (D.) immutatus sp. nov. (Laos), O. (D.)
buccatus sp. nov. (Laos), O. (D.) innexus sp. nov.
(Borneo), O. (D.) furviceps sp. nov. (Borneo), O. (D.)
compressus sp. nov. (Borneo), O. (D.) bituberculatus sp.
nov. (Borneo),
O. (D.) tensicornis sp. nov. (Borneo),
O. (D.) rogans sp. nov. (Borneo), O. (D.)
tuberifer sp. nov. (Borneo), O. (D.) bulbifer sp. nov.
(Sulawesi Tengah), O. (D.) bulbosus sp. nov. (Philippines:
Mindanao), and O. (D.) hirsutus sp. nov. (Borneo);
Amaurodera dentissima sp. nov. (Myanmar), A. setosicollis sp. nov.
(Borneo), and A. floreni sp. nov. (Borneo); Drusilla trituberculata
sp. nov. (China: Hubei), D. longispinosa sp. nov. (Borneo), D. iniqua
sp. nov. (Borneo),
and D. samalica sp. nov. (Philippines: Mindanao); Witteia arboris
sp. nov. (Borneo);
Tensusa procera sp. nov. (Laos);
Pheidologitonetes quadriplicatus sp. nov. (Laos), P. artiplicatus
sp. nov. (Laos), P. acer sp. nov. (Vietnam), P. retunsus sp. nov.
(Vietnam), P. sagittatus sp. nov. (Laos), and P. punctatus sp.
nov. (Laos); Zyras (Zyras) bituberosus sp. nov. (Laos), Z.
(Z.) semicalvus sp. nov. (Laos), Z. (Z.)
janetlarae sp. nov. (Borneo), Z. (Z.) parvihirtus sp.
nov. (Borneo), Z. (Z.) cyaniceps sp. nov. (Irian
Jaya), Z. (Z.) maindai sp. nov. (Irian Jaya), Z. (Z.)
interruptus sp. nov. (Irian
Jaya), Z. (incertae sedis) pressitibialis sp. nov. (Borneo), and Z.
(i. s.) cuneatus sp. nov. (Laos); Mimaenictus bisetosus sp. nov.
(Thailand); Scapaenictus longiscapus sp. nov. (Laos); Planusa
granulosa sp. nov. (Laos); Sulciusa boops sp. nov. (Borneo). Four
monotypical genera are described: Tensusa gen. nov. (type species: T.
procera); Scapaenictus gen. nov. (type species: S. longiscapus);
Planusa gen. nov. (type species: P. granulosa); Sulciusa
gen. nov. (type species: S. boops). Nine new combinations are
established: Orphnebius mirabilis (Pace, 2014), comb. nov. (ex Keratodegnathus
Pace, 2014); Pheidologitonetes thaifuscicollis (Pace, 2012), comb. nov. (ex
Drusilla Leach,
1819); P. rubricollis (Cameron, 1939), comb. nov. (ex Episkiodrusilla Pace,
2013); P. malayrubricollis (Pace, 2013), comb. nov. (ex Episkiodrusilla);
P. neocoenonicacollis (Pace, 2008), comb. nov. (ex Episkiodrusilla);
P. vedda Pace, 1992, comb. nov. (ex Drusilla); P. veluticollis
(Pace, 2013) , comb. nov. (ex Episkiodrusilla); Drusilla nilgiriensis
(Pace, 2001), comb. nov. (ex Pheidologitonetes); D. bartolozzii (Pace,
2001), comb. nov. (ex
Pheidologitonetes). Four new species groups are proposed within Orphnebius
Motschulsky, 1858: the O. mollis, O. colitus, and O. tensus
subgroups of the O. hauseri group, and the O. hirsutus species
group of Deroleptus Bernhauer, 1915. The intrageneric affiliations of Orphnebius,
their systematic implications, and their nomenclatural consequences are
discussed. Two synonymies are established: Deroleptus Bernhauer, 1915 = Aulacothoracobius
Bernhauer, 1929, syn. nov.; Pheidologitonetes
Cameron, 1939 = Episkiodrusilla Pace, 2013, syn. nov. Updated
catalogues of Orphnebius sensu lato and of Pheidologitonetes are provided. Additional records of 59 previously
described species are reported, among them numerous new country records.
587:
The
Cephennium fauna of Turkey and the Middle East (Coleoptera:
Staphylinidae: Scydmaeninae).
Only
two species of Cephennium Mόller & Kunze, 1822 were previously
known from Turkey and the Middle East including Lebanon, Israel, Jordan, Iraq,
and Iran. An examination of material collected during numerous field trips to
Turkey and Jordan in the past four decades revealed that the Cephennium
fauna of Turkey is remarkably diverse. In total, 25 named species are
recognized, diagnosed, illustrated, and assigned to four species groups. Based
on zoogeographic, morphological, and additional evidence, C. bodemeyeri Reitter,
1903, previously a subspecies of C. perispinctum Kolenati, 1846, is elevated to
species rank. Twenty-three species are described for the first time: Cephennium
fractum spec. nov. (Turkey: Bursa, Balıkesir, Kocaeli, Ηanakkale), C.
infractum spec. nov. (Turkey:
Bursa), and C. immutatum spec.
nov. (Turkey: Izmir, Muğla, Adıyaman) of the C. perispinctum
group; C. ericinum spec. nov.
(Turkey: Ordu), C. varispinosum spec.
nov. (Turkey: Ordu), C. victrix spec.
nov. (Turkey: Sinop), C. spinigerum spec.
nov. (Turkey: Bartin), and C. digitale spec.
nov. (Turkey: Zonguldak) of the C. ericinum group; C. reductum spec.
nov. (Turkey: Konya, Antalya, Mersin, Kastamonu) of the C. jonicum group;
C. corruptum spec. nov. (Turkey: Mersin, Adana), C.
marasicum spec. nov. (Turkey: Kahramanmaraş), C.
dolucanum spec. nov. (Turkey: Kahramanmaraş), C. fimbriatum spec. nov. (Turkey: Adana), C. besucheti spec.
nov. (Turkey: Kahramanmaraş), C.
affimbriatum spec. nov. (Turkey: Kahramanmaraş), C. kartalicum spec. nov. (Turkey: Gaziantep), Cephennium nemruticum spec. nov. (Turkey: Adıyaman), C.
unguicum spec. nov. (Turkey: Aydın), C. deplectens spec.
nov. (Turkey:
Izmir, Manisa), C. sicaferum spec. nov. (Turkey:
Aydın), C. geminum spec.
nov. (Turkey: Antalya), C.
duplum spec. nov. (Turkey: Muğla), and C. jordanicum spec. nov. (Northwest Jordan) of the C.
corruptum group. Several additional species, all of them represented
exclusively by females and most likely belonging to the C. corruptum
group, remain undescribed. A key to the named species of Turkey and the Middle
East is provided. The distributions of the species recorded from Turkey are
mapped. In an appendix, C. liguliferum spec. nov. (Southeast Bulgaria), a
close relative of C. bodemeyeri and C. perispinctum, is described
and illustrated.
588:
New
species, synonymies, combinations, and records of micropterous Athetini from
China (Coleoptera: Staphylinidae: Aleocharinae).
Two
micropterous and locally endemic species of Bellatheta Roubal,
1928 and four of Atheta Thomson,
1858, subgenus Microdota Mulsant
& Rey, 1873, from China are described and illustrated: Bellatheta
qiliana spec. nov. (Qinghai, Gansu); B. aucticeps spec. nov. (Yunnan:
Gaoligong Shan); Atheta (Microdota) clarata spec. nov. (Yunnan:
Gaoligong Shan); A. (M.) minica spec. nov. (Sichuan:
Songpan env.); A. (M.) qinlingica spec. nov. (Shaanxi:
Qinling Shan); A. (M.) biformis spec. nov. (Gaoligong
Shan). New illustrations are provided for two previous described Microdota
species. One genus-group and one species-group synonymy are proposed: Bellatheta
Roubal, 1928 = Oroekklina Pace,
1999, syn. nov.; Atheta (Microdota)
amischoides Assing, 2006 = A
(M.) wuliangensis Pace,
2017, syn. nov. Five new combinations are proposed: Bellatheta daxuensis
(Pace, 1999), comb. nov., B. excaecata (Assing,
2009), comb. nov., B. proiecta (Assing,
2002), comb. nov., B. smetanai (Pace, 2004), comb. nov. (all of them previously in Oroekklina),
and Franzidota geostiboides (Assing,
2011), comb. nov. (ex Platyola Mulsant
& Rey, 1875). Additional records of two species of Bellatheta
and of five species of Microdota are reported from China. Catalogues of
the Bellatheta species and the micropterous Microdota species
recorded from China are provided. Including the new species and synonymies, 15
species of Bellatheta, seven of them micropterous, and 85 of Microdota,
19 of them micropterous, have been reported from China. The Microdota
fauna of Taiwan is composed of 13 species, four of them micropterous. The
distributions of the micropterous Bellatheta species of China are mapped.
590:
A
revision of Palaearctic, Oriental, and New Guinean Pseudolathra. VI. Four
new species, two new synonymies, and additional records (Coleoptera,
Staphylinidae, Paederinae).
Four
species of Pseudolathra Casey, 1905
from Nepal and Western New Guinea are described and illustrated: P.
brevalata nov.sp. (Nepal); P. maindai nov.sp. (Western New Guinea:
Foja Mountains); P. atriflava nov.sp. (Western New Guinea: Foja
Mountains); P. brevirimata nov.sp. (Western New Guinea: Foja Mountains).
Except for the latter species, all of them belong to the P. nigerrima
group. Two synonymies are proposed: Pseudolathra
quadricollis (Fauvel, 1875) = Lathrobium
boyadjiani Rambousek, 1907,
nov.syn.; Pseudolathra himalayana Assing,
2012 = P. gansuensis Li, Cai &
Zhou, 2021, nov.syn. Additional records of eight previously described are
reported from the Palaearctic and Oriental regions, as well as from New Guinea.
591:
On
the taxonomy and zoogeography of Stilicoderus and Stiliderus VII.
Two new species from the Oriental region, a revalidation, and additional records
(Coleoptera: Staphylinidae: Paederinae).
Two
species of Stilicoderus Sharp, 1889 are described and illustrated: Stilicoderus
opupifer sp. nov. (North Vietnam) of the S. japonicus group and S.
colitus sp. nov. (Thailand, Laos) of the S. discalis group, a species
previously confounded with S. discalis Fauvel,
1895. Stilicoderus subseriatus (Eppelsheim, 1895) from Myanmar,
previously a junior synonym of S. discalis, is revalidated, diagnosed,
and illustrated. A lectotype is designated for Stilicus subseriatus
Eppelsheim, 1895. Additional records of 13 species of Stilicoderus and of
seven species of Stiliderus Motschulsky, 1858 are reported, among them
four new country records from Laos (two), Myanmar (one), and Nepal (one). Stilicoderus
now includes 116, Stiliderus 51 species.
592:
New
species of Staphylinidae from Pakistan (Coleoptera: Staphylinidae: Aleocharinae,
Paederinae, Pselaphinae).
Six
species of Staphylinidae from North Pakistan are described and illustrated: Cousya
prominens Assing sp. nov. (Punjab),
Tachyusa semigilva Assing sp. nov. (Punjab),
Platyola lippa Assing sp. nov. (Punjab), and P. invidens Assing
sp. nov. (North-West Frontier) of the Aleocharinae, Micranops viti Assing
sp. nov. of the Paederinae, and Euplectus pakistanicus Brachat sp.
nov. of the Pselaphinae.
593:
On
the taxonomy and zoogeography of the Caucasian genus Pseudotyphlopasilia
(Coleoptera: Staphylinidae: Aleocharinae).
A
study of recently collected material of Pseudotyphlopasilia Pace, 1983
revealed that the genus is confined to the Caucasus region (West Georgia;
Russian part of West Caucasus) and includes ten species, all of them
anophthalmous, micropterous, and locally endemic. Typhlusa Pace, 1983,
previously a subgenus of Pseudotyphlopasilia and including only a single
species from Montenegro, is attributed generic rank. Primarily based on the
structure of the median lobe of the aedeagus and the shape and chaetotaxy of the
paramere, Pseudotyphlopasilia is removed from the Athetini and
tentatively placed in the Meoticina of the Oxypodini. Eight of the Pseudotyphlopasilia
species are described for the first time: P. acris sp. nov. (North
Georgia: Svaneti); P. angulata sp. nov. (Southwest Georgia: estreme west
of Meskheti range); P. baculata sp. nov. (Southwest Georgia: western
Meskheti and Shavsheti ranges); P. carolae sp. nov. (Southwest Georgia:
western Shavsheti range); P. depressa sp. nov. (North Georgia:
Mtskheta-Mtianeti); P. kakhetica sp. nov. (North Georgia: Kakheti
region); P. kociani sp. nov. (North Georgia: Tusheti region); P.
vespertina sp. nov. (Russia: Krasnodar region). Pseudotyphlopasilia
cavernicola (Assing, 2007), previously a synonym of P. coeca
(Eppelsheim, 1878), is revalidated. A lectotype is designated for Leptusa
coeca Eppelsheim, 1878. The currently known distributions of the species are
mapped. The morphological adaptations, the methods with which they were
collected, and the habitats where they were found suggest that Pseudotyphlopasilia
species inhabit deep litter layers and the mineral soil of moist, mostly
deciduous forests.
594:
On
the Silusa fauna of China (Coleoptera: Staphylinidae: Aleocharinae:
Silusini).
Four
species of Silusa Erichson, 1837 are described and illustrated: S.
edentula sp. nov. (China: Sichuan: Emei Shan); S. triangularis sp.
nov. (China: Yunnan: Gaoligong Shan); S. mastigans sp. nov. (China:
Sichuan: Gongga Shan); S. turgida sp. nov. (China: Sichuan: Gongga Shan).
The two species from Gongga Shan are micropterous. Additional records of four
previously described species are reported; the male sexual characters of three
of them are illustrated. A catalogue of the Silusa species recorded from
China is provided. The Silusa fauna of China is now represented by 14
species with three species from Sichuan micropterous, that of the Palaearctic
region includes 31 species.
595:
Review
of Palaearctic Autalia VIII. A new species from Sichuan, China, and
additional records (Coleoptera: Staphylinidae: Aleocharinae).
Autalia
tibetana
sp. nov. (China:
Sichuan: Ganzi Tibetian Autonomous Prefecture), a winged species found in marmot
burrows at an altitude of 4200 m, is described and illustrated. Additional
records of seven species of Autalia Leach, 1819 are reported from the
Palaearctic region, among them a first record from Georgia and a new province
record from China. The genus is now represented in the Palaearctic region by 18
species (plus one nomen dubium).
596:
A
new genus and species of Oxypodini from Sichuan, China (Coleoptera:
Staphylinidae: Aleocharinae).
Sinolophus
gen. nov. of the Oxypodini (tentative subtribal assignment: Oxypodina) and its
type species Sinolophus rubellus sp. nov. (China: Sichuan) are described
and illustrated. The new genus combines a remarkable mix of characters of Eurylophus
J. Sahlberg, 1876 on the one hand and of Echidnoglossa Wollaston, 1864
and allied genera on the other.
597:
Two
new species of Phymatura from China (Coleoptera: Staphylinidae:
Aleocharinae).
FPhymatura
angulata
sp. nov. (China:
Yunnan) and P. russa sp. nov. (China: Shaanxi) are described and
illustrated. Phymatura cooteri Assing, 2005 is reported from Japan for
the first time. Including the new species, Phymatura J. Sahlberg, 1876 is
represented in the Palaearctic region by 14 species, at least one of them of
doubtful generic assignment.
598:
The
first record of the myrmecophilous genus Thiasophila from China
(Coleoptera: Staphylinidae: Aleocharinae).
Thiasophila
gansuica
sp. nov. (South
Gansu), the first representative of the genus to be recorded from China, is
described, illustrated, and distinguished from other congeners recorded from the
East Palaearctic region. The myrmecophilous species is associated with Formica
chinensis Wheeler, 1913 of
the F. truncorum group. Thiasophila Kraatz, 1856 currently
includes 18 species worldwide, 13 of which are distributed in the Palaearctic
region.
599:
On
the taxonomy, diversity, and ecology of the Amischa Thomson,
1858 species of the Palaearctic Region
(Coleoptera: Staphylinidae: Aleocharinae).
Based
on a revision of types and additional material, the genus Amischa Thomson,
1858 is represented in the Palaearctic region by sixteen species, i.e., little
more than half the species number previously recorded from this region (30
species). Unlike many other genera of Athetini, the genus is characterised by
several pronounced synapomorphies. The species are assigned to two species
groups, the monotypical A. puetzi and the A. analis groups (15
species); several distinct lineages are identified within the A. analis
group. Diagnoses and illustrations are provided for all the species recorded
from the Palaearctic region. Four species are described for the first time: Amischa
iberica sp.n. (South Spain); A. inexcisa sp.n. (Turkey, West
Caucasus, Georgia, Hungary); A. truncata sp.n. (South Turkey); A.
discolor sp.n. (Kyrgyzstan, Kazakhstan). Nine synonymies are proposed: Amischa
bifoveolata (Mannerheim,
1830) = A. filum (Mulsant
& Rey, 1870), syn.n., =
A. minima Mulsant
& Rey, 1873, syn.n., =
A. uhligi Pace,
1987; syn.n.; A. decipiens (Sharp,
1869) = A. simplex Benick,
1982, syn.n.; A.
kashmirica Cameron,
1939 = A. sporadica Benick,
1967, syn.n.; A. nigrofusca (Stephens,
1832) = A. jugorum Scheerpeltz,
1956, syn.n.; A.
puetzi Assing,
2005 =
A. sichuanensis Pace,
2011, syn.n.; A.
setifera Benick,
1982 = A. andreasi Muona,
1990, syn.n., =
A. rougemonti Pace,
1998; syn.n. Records previously reported as A. filum refer to four
species, two of them newly described: A. bifoveolata, A. kashmirica,
A. inexcisa, A. truncata. Eight species described in or previously
moved to Amischa are transferred to Atheta Thomson,
1858: Atheta
(Microdota)
pioncolyata (Pace,
1991), comb.n.; Atheta (M.)
nana (Pace,
1998), comb.n.; Atheta
(Microdota?)
phallosetigera (Pace,
1986), comb.n.; Atheta coprophila (Pace,
1991), comb.n.; Atheta
dhaulagiriensis (Pace,
2015), a secondary junior homonym replaced with Atheta dhaulagirica
nom.n.; Atheta fraudulenta Pace,
2006, comb.n.;
Atheta (Mocyta)
sahlbergi (Eppelsheim,
1893), comb.n.;
Atheta niponensis (Sharp,
1888), comb.n.
Lectotypes are designated for Homalota filum Mulsant
& Rey, 1870, Amischa strupii Scheerpeltz,
1967, and Homalota leporina Fauvel,
1881. Based on extensive field studies using pitfall traps, soil extractions,
and photo-eclectors, and conducted in Niedersachsen (North Germany) from 1981 to
2001, the sex ratios, life histories, seasonal density and epigeic activity
dynamics, and parasitism of four species are assessed and discussed. Amischa
analis (Gravenhorst,
1802) is particularly remarkable in that this species is parthenogenetic
and macropterous across most its range, while scattered bisexual and
brachypterous populations are confined to high-altitude habitats in the South
Alps and mountain ranges in the northern Iberian Peninsula. Amischa nigrofusca
(Stephens,
1832) and A. decipiens (Sharp,
1869) are regionally parthenogenetic, with males significantly rarer also in
bisexual populations. A catalogue of the Amischa species represented in
the Palaearctic region is provided. Numerous new country records are reported.
The distributions of four species are mapped.
600:
Zur
Carabiden- und Staphylinidenfauna von Kies- und Schotterufern im Nord- und
Westharz, Sόdniedersachsen
(Coleoptera).
A
study of gravel banks of rivers and streams conducted in the northern and
western parts of the Harz mountains, southern Niedersachsen, in spring and
summer 2020 yielded 5122 adult specimens belonging to 251 species of Carabidae
(1955 specimens; 69 species) and Staphylinidae (3167 specimens; 182 species). As
many as nine species, one of Carabidae and eight of Staphylinidae, were recorded
from the region for the first time; twelve additional species of Staphylinidae
had not been recorded from the area for decades, two have them for more than a
century. The most remarkable records are discussed regarding their ecological
and faunistic significance.
601:
The
Afropselaphus fauna of South Turkey and the Middle East (Coleoptera,
Staphylinidae, Pselaphinae).
Only
a single named species of Afropselaphus Jeannel,
1950, A. clavigeroides (Reitter,
1882) from Lebanon, was previously known from the region comprising South Turkey
and the Middle East. Based on a study of material collected since the 1980s, 18
additional species are described and illustrated, all of them micropterous and
locally endemic: Afropselaphus muglanus nov.sp. (Southwest Turkey: Muğla);
A. pisidicus nov.sp. (Southwest Turkey: western Antalya); A. wolfi
nov.sp. (South Turkey: eastern Antalya); A. sinuatus nov.sp. (South
Turkey: eastern Antalya); A. tenuispinosus nov.sp. (South Turkey:
northern Antalya); A. frederici nov.sp. (South Turkey: western Mersin);
A. cilicicus nov.sp. (South Turkey: Mersin); A. adanicus nov.sp.
(South Turkey: Adana); A. longispinosus nov.sp. (South Turkey:
Kahramanmaraş, Osmaniye); A. spoliatus nov.sp. (South Turkey:
Osmaniye); A. latibasalis nov.sp. (South Turkey: Osmaniye); A.
trispinosus nov.sp. (South Turkey: Osmaniye); A. kizilicus nov.sp.
(South Turkey: Hatay); A. bispinosus nov.sp. (South Turkey: Hatay); A.
meybohmi nov.sp. (Jordan); A. inaequalis nov.sp. (North Israel: Upper
Galilee); A. aequalis nov.sp. (North Israel: Upper Galilee); A.
assmanni nov.sp. (North Israel: Upper Galilee). All the species are assigned
to the newly established A. clavigeroides group, which additionally
includes four species from Crete. A catalogue of the Afropselaphus
species represented in South Turkey and the Middle East is provided; their
distributions are mapped. The genus now includes a total of 45 species
distributed in the West Palaearctic region eastwards to Uzbekistan.
602:
A
new subgenus and a remarkable new species of Domene from South China,
with additional records of the genus from China, Taiwan, and Vietnam
(Coleoptera: Staphylinidae: Paederinae).
Domene
hybrida
spec. nov. (China:
Guizhou) is described, illustrated, and distinguished from other species of the
genus known from China. The species is peculiar in that it combines
morphological characters found in Domene Fauvel,
1873 with characters observed in other genera of Lathrobiina, particularly Lobrathium
Mulsant & Rey, 1878 and Pseudobium
Mulsant & Rey, 1878. Unlike all
other congeners previously recorded from China, it does not belong to the
subgenus Macromene Coiffait,
1982, which is why it is assigned to a subgenus of its own, Lobramene
subgen. nov. Additional records of six Domene species are reported from
China, Taiwan, and North Vietnam. Including two species of doubtful generic
affiliations, the genus now includes 86 named species, 22 of which have been
recorded from China, three from Taiwan, and two from Vietnam.
603:
On
the Lathrobiina of Taiwan II. A new species and an additional record of Lobrathium
(Coleoptera: Staphylinidae: Paederinae).
A
micropterous species of Lobrathium Mulsant & Rey, 1878 of the L. stimulans
group from Taiwan is described and illustrated: L. colligatum. spec. nov.
(Hualien Hsien: Taroko National Park). An additional record of L. bilobatum
Assing,
2010 is reported. The subtribe Lathrobiina is now represented in Taiwan by 44
species.
604:
A
revision of Othiini XXII. A new species, a new synonymy, and additional records
of Othius from Northwest Georgia (Coleoptera: Staphylinidae:
Staphylininae).
A
total of 373 specimens of Othius Stephens,
1829 was collected during three field trips to West Georgia conducted in June,
July/August, and October 2021. The material is represented by eight species. One
of them is described and illustrated: Othius
egrisicus spec. nov. (Georgia: Samegrelo-Zemo Svaneti), a representative of
the O. crassus group. The following synonymy is proposed: Othius hebes
Assing & Solodovnikov, 1998 = O.
fastigatus Assing & Solodovnikov,
1998, syn. nov. Two species are recorded for the first time since their
respective original descriptions. The distributions of several species endemic
in Northwest Georgia are clarified and mapped. The Georgian Othius fauna
is currently represented by 16 described species, eleven of them endemic, and
the genus as whole currently includes 136 species and subspecies.
605:
A
revision of Medon. XII. Two new
species and a review of the fauna of Georgia (Coleoptera: Staphylinidae:
Paederinae).
Two
species of Medon Stephens,
1833 are described and illustrated: M. steggiae spec. nov. (Northwest
Georgia: Samegrelo-Zemo Svaneti, Abkhazia) of the M. fusculus group and M.
bisinuatus spec. nov. (China: Zhejiang) of the M. profundus group. A
revision of previously examined and recently collected material from Georgia
revealed that the fauna of this country currently includes six species and that
some of the previous records were based on misidentification. All the known
records of the genus from Georgia are mapped.
606:
New
species and records of Leptusa from the Georgian Caucasus (Coleoptera:
Staphylinidae: Aleocharinae).
Three
field trips to West Georgia conducted in 2021 yielded 923 specimens of Leptusa
Kraatz, 1858 belonging to ten
species, two of which are described and illustrated: Leptusa (Neopisalia)
pugiofera spec. nov. (Georgia:
Kvemo Svaneti); L. (N.)
insculpta spec. nov. (Georgia: Zemo Svaneti). The
distributions of five species are mapped. The Leptusa fauna of the
Caucasus region sensu lato is now represented by a total of 40 species, 21 of
which (11 exclusive) have been recorded from Georgia. The Palaearctic fauna
includes 420 species and 74 subspecies in 71 subgenera.
607:
Revision
of Geostiba of the West Palaearctic
region. XXVIII. New species and records from the Georgian Caucasus (Coleoptera:
Staphylinidae: Aleocharinae).
A
study of 485 specimens of Geostiba Thomson,
1858, 482 of which were collected during three field trips to West Georgia
conducted in 2021, yielded eleven species, two of which are described and
illustrated: Geostiba (Sibiota) brevifistula spec. nov. (Georgia:
Kvemo Svaneti) and G. (S.)
egrisica spec. nov. (Georgia: Samegrelo-Zemo Svaneti). Both
species belong to the G. bituberculata group. Geostiba (Geostiba)
sororcula Assing, 2001, a species previously known only from Northeast
Turkey and Armenia, is reported from Georgia for the first time. The
distributions of five species are mapped. The Geostiba fauna of the
Caucasus region sensu lato is now represented by a total of 50 species, 30 of
which belong to the subgenus Sibiota Casey,
1906.
608:
On
the Nanophthalmus fauna of Georgia (Coleoptera: Staphylinidae:
Scydmaeninae).
Only
two named species of Nanophthalmus Motschulsky, 1851 were previously known from
Georgia. A study of mostly recently collected material from Georgia yielded nine
additional species, one of them previously described and reported from Georgian
territory for the first time, two unnamed (male unknown), and six newly
described and illustrated: Nanophtalmus egrisicus spec. nov. (Zemo
Svaneti); N. svaneticus spec. nov. (Zemo Svaneti); N. truncatus
spec. nov. (Racha); N. brachati spec. nov. (Kvemo Svaneti); N.
convexus spec. nov. (Adjara);
N. simplex spec. nov. (Adjara).
The identity and taxonomic status of N. armeniacus (Reitter,
1884), a previously probably misinterpreted species, are discussed. The Georgian
Nanophthalmus species are anophthalmous and locally endemic, three in the
Lesser Caucasus and five in the Greater Caucasus west of South Ossetia. Their
distributions are mapped. The genus now includes a total of 19 described
species.
609:
New
records, detailed distribution and abundance of rove-beetles (Insecta,
Coleoptera, Staphylinidae) collected between 1990 and 2015 in Azores (Portugal)
with an updated checklist.
The dataset we present consists of an
inventory compiling all records and knowledge about Staphylinidae (Insecta,
Coleoptera) in the Azores and is part of a long-term monitoring performed
between 1990 and 2015 in different habitat types of eight islands of the Azores
Archipelago. Most samples come from the BALA project (Biodiversity of Arthropods
from the Laurisilva of Azores) that sampled native forests in the Azores.
Additional sampled habitats include exotic forests, intensive and semi-natural
pasturelands, orchards, caves and lava flows. Most of the records (about 96.7%)
were collected in standardised sampling campaigns, which included pitfall traps
and beating transect protocols. Non-standardised records are based on
hand-collecting and sifting, as well as cave, colour and malaise traps. We
provide a long-term inventory of Staphylinidae (Insecta, Coleoptera) collected
in the course of several standardised sampling campaigns and recorded with
non-standardised methods. We collected a total of 10744 specimens belonging to
69 identified species of Staphylinidae, which represents 51% of the species
known from the Azores Archipelago.
Four endemic species were sampled, representing 40% of the known Azorean endemic
species. From this dataset, seven species are new for the Azores: Aleochara
funebris Wollaston, 1864; Amischa
forcipata Mulsant & Rey, 1873; Bledius
unicornis (Germar,
1825); Carpelimus troglodytes (Erichson,
1840); Cypha seminulum (Erichson,
1839); Paraphloeostiba gayndahensis (MacLeay,
1871); Tachyporus caucasicus Kolenati,
1846. We also registered a total of 66 new island records for eight Azorean
islands. This
contribution continues a series of publications on the distribution and
abundance of Azorean arthropods. We also provide an updated list of Azorean
rove-beetles (Staphylinidae) that now includes 136 species, ten of them
considered Azorean endemics.
610:
On
the taxonomy and zoogeography of Paederus. VI. Two new species from Nepal
and new records from the Palaearctic and Oriental regions (Coleoptera:
Staphylinidae: Paederinae).
Two
micropterous species of Paederus Fabricius,
1775 from East Nepal are described and illustrated: Paederus (incertae
sedis) digitalis spec. nov. and P. (incertae sedis) acifer
spec. nov. They are distinguished from other micropterous and geographically
close congeners. Including the new species, the Paederus fauna of Nepal
currently includes 37 species, 24 of them exclusive. Additional records of two
widespread species are reported from the Chinese province Yunnan and from
Vietnam.
611:
On
the taxonomy and zoogeography of the Caucasian genus Pseudotyphlopasilia
(Coleoptera: Staphylinidae: Aleocharinae). IV. Two new species and additional
records from Northwest Georgia.
Two
field trips to Northwest Georgia conducted in 2021 yielded 29 specimens of the
Caucasian genus Pseudotyphlopasilia
Pace,
1983, most of them collected by soil-washing. The material belongs to five
species, two of them described and three of them unnamed. Two of the unnamed
species are described and illustrated: P. confusa spec. nov. (Racha:
environs of Lailashi) and P.
egrisica
spec. nov. (Zemo
Svaneti: Egrisi Range). One
species remains unnamed for want of males. Additional records are provided for
two previously described species, including Pseudotyphlopasilia coeca (Eppelsheim,
1878) (type species of the genus) of which only few and mostly old specimens
were previously available. New illustrations are provided for P. coeca
and P. acris Assing,
2021. The genus currently contains a total of twelve named species, all of them
anophthalmous and micropterous, with eleven of them distributed in Georgia and
one in the Russian West Caucasus. The distributions of five species are mapped.
612:
On
Orsunius V. Seven new species from Borneo and Vietnam (Coleoptera:
Staphylinidae: Paederinae: Medonina).
Seven
species of Orsunius Assing, 2011
are described and illustrated, six of them from Borneo and one from Vietnam: Orsunius
weigeli spec. nov. (North Vietnam), O. floreni spec. nov. (Malaysia:
Sabah), O. flavoniger spec. nov. (Malaysia: Sabah), O. incitatus
spec. nov. (Malaysia: Sabah), O. curvicollis spec. nov. (Malaysia:
Sabah), O. tricolor spec. nov. (Malaysia:
Sabah), and O. arboris spec. nov. (Malaysia: Sabah). The genus now
includes 31 named extant species distributed in the southern East Palaearctic,
the Oriental, and the northern Australian regions.
613:
A
new species of Acrolocha from North Iran (Coleoptera: Staphylinidae:
Omaliinae).
A new
species of the Holarctic omaliine genus Acrolocha Thomson,
1858 from Gilan province, Northwest Iran, is described, illustrated, and
distinguished from other West Palaearctic congeners. The genus now includes a
total of 16 species, seven of which are distributed in the West Palaearctic
region.
614:
Two
new species of Bergrothia Reitter,
1884, with a review of the Bergrothia fauna of Georgia (Coleoptera:
Staphylinidae: Pselaphinae).
Two
species of Bergrothia Reitter,
1884, a small Caucasian genus of Amauropini, are described and illustrated: B.
svanetica spec. nov. (Georgia: Svaneti region) and B. simplex spec.
nov. (Northeast Turkey: Ordu, Giresun). Including the new species, the genus now
includes a total of nine species distributed in Georgia (six species), Northeast
Turkey (three species, and Azerbaijan (one species). Based on abundant material
collected during seven field trips conducted to Georgia between 2016 and 2021,
the partly allo- and partly sympatric distributions of the Bergrothia
species in Georgia are clarified and mapped. All of them are confined to the
west of the country.
615:
The
first records of the genus Tychobythinus Ganglbauer,
1896 from Georgia (Coleoptera: Staphylinidae: Pselaphinae).
Three
microphthalmous and presumably locally endemic species of Tychobythinus Ganglbauer,
1896, a pselaphine genus previously unknown from Georgia, are described
and illustrated: T. meskheticus spec. nov. (Southwest Georgia: Meskheti
Range), T. eximius spec. nov. (Northwest Georgia: Egrisi Range), and T.
egrisicus spec. nov. (Northwest Georgia: Egrisi Range). Tychobythinus
eximius is particularly remarkable in that it is subject to a unique sexual
dimorphism of the antennae and the pronotum. Including the new species, Tychobythinus
is now represented in the Palaearctic region by 95 named species and two
subspecies.
616:
Monograph
of Palaearctic, Oriental, and New Guinean Pinophilina (Coleoptera:
Staphylinidae: Paederinae).
The
Palaearctic, Oriental, and New Guinean fauna of the Pinophilina was previously
represented by 78 valid species and one subspecies in seven genera, with 68 (87
%) of the species in the nominal genus Pinophilus Gravenhorst, 1802. This
paederine subtribe had never been subject to a comprehensive revision. In order
to assess the taxonomy, systematics, and diversity of the Pinophilina in the
study region, the accessible type material of previously named species, as well
as numerous additional specimens, both previously identified and unidentified,
from the major public and private collections were examined. In addition,
relevant genera and species, in particular Pinophilus (type genus of
Pinophilina) and Pinophilus latipes Gravenhorst, 1802 (type species of Pinophilus),
from other zoogeographic regions were included in the study. The revision
revealed that Pinophilus is absent from the Palaearctic and Oriental
regions and probably confined to the New World. The subtribe is represented in
the Palaearctic and Oriental regions, including New Guinea, by as many as 309
named species in 16 genera, and none of the previously described species remains
in Pinophilus. Numerous species groups are newly established particularly
in speciose genera. Except for two recently described species and one species
whose type material is inaccessible, all the species are (re-)described and
illustrated.
Eleven genera and 241 species are described for the first time: Mimopinophilus
compar sp. nov. (Iran; Oman; Pakistan; India); M. hamatus sp. nov.
(Indonesia: Sulawesi); M. triangularis sp. nov. (South India; Sri Lanka);
M. uncinatus sp. nov. (Indonesia: Sumbawa); Neopinophilus abscisus
sp. nov. (Malaysia: Selangor); N. crockermontis sp. nov. (Malaysia:
Sabah); N. edentulus sp. nov. (South India); N. fulgens sp. nov.
(Malaysia: Sarawak); N. glaber sp. nov. (Malaysia: Sarawak); N.
kinabaluicus sp. nov. (Malaysia: Sabah); N. mindoronus sp. nov. (Philippines:
Mindoro); N. muluanus sp. nov. (Malaysia: Sarawak); N. palawanus
sp. nov. (Philippines: Palawan); N. rubellus sp. nov. (Indonesia:
Sumatra); N. sandakanus sp. nov. (Malaysia: Sabah); N. selatanus
sp. nov. (Indonesia: Sulawesi); N. tensicollis sp. nov. (Indonesia:
Sulawesi); N. umbilicatus sp. nov. (Sri Lanka); Phinopilus acutior
sp. nov. (China: Sichuan, Zhejiang); P. adclinis sp. nov. (China: Yunnan;
Laos; Vietnam); P. adesus sp. nov. (Papua
New Guinea); P. alishanus sp. nov. (Taiwan); P. ambangus sp. nov.
(Indonesia: Sulawesi); P. ambulans sp. nov. (Philippines: Leyte); P.
ampliatus sp. nov. (Papua New Guinea); P. amplicornis sp. nov. (North
India); P. angustior sp. nov. (Indonesia:
Kalimantan Selatan); P. arcticornis sp. nov. (Papua
New Guinea); P. arquatus sp. nov. (Indonesia:
Java); P. artior sp. nov. (China:
Fujian, Zhejiang); P. auriculatus sp. nov. (North India); P. aviformis
sp. nov. (Singapore); P. baculatus sp. nov. (Malaysia;
Indonesia: Sumatra); P. biacifer sp. nov. (Sri Lanka); P. bilamellatus
sp. nov. (China: Sichuan); P. boops sp. nov. (Malaysia: Sarawak); P.
brevior sp. nov. (Malaysia:
Sarawak); P. cacharicus sp. nov. (Northeast India); P. cacuminatus sp.
nov. (Thailand; Laos; Cambodia); P. callicollis sp. nov. (Papua New
Guinea, New Ireland, New Britain; Indonesia: Maluku Islands); P. candens sp.
nov. (Malaysia;
Indonesia); P. caninus sp. nov. (Thailand;
Laos); P. celebesicus sp. nov. (Indonesia:
Sulawesi); P. cernens sp. nov. (Malaysia: Sabah); P. chiangicus
sp. nov. (Thailand);
P. clavifer sp. nov. (Malaysia: Sabah); P. clinatus sp. nov. (New
Guinea); P. coauditus sp. nov. (Thailand); P. collevatus sp. nov.
(Western New Guinea; Papua New Guinea); P. confossus sp. nov.
(Philippines: Palawan); P. cordiformis sp. nov. (South India); P.
crenatus sp. nov. (Indonesia: Western New Guinea); P. cubans sp. nov.
(Malaysia:
Sabah); P. cultellatus sp. nov. (Brunei);
P. curtatus sp. nov. (South India); P. digitalis sp. nov. (Cambodia);
P. edolatus sp. nov. (Indonesia:
Western New Guinea); P. eminens sp. nov. (Malaysia: Peninsular, Sabah,
Sarawak); P. esuriens sp. nov. (Malaysia: Sarawak); P. excavatus
sp. nov. (Cambodia); P. fibulatus sp. nov. (Thailand); P. figens
sp. nov. (Indonesia:
Java); P. firmicornis sp. nov. (Malaysia);
P. fodens sp. nov. (Thailand); P. forticornis sp. nov. (Indonesia:
Sumatra); P. garonus sp. nov. (India:
Meghalaya); P. gradiens sp. nov. (Indonesia: North Maluku); P.
grossicornis sp. nov. (Malaysia);
P. hainanus sp. nov. (China:
Hainan); P. hastatus sp. nov. (Vietnam); P. hebes sp. nov.
(Malaysia: Sarawak; Brunei); P. inspicatus sp. nov. (Nepal; India;
Myanmar; China; Thailand; Laos; Vietnam; Indonesia: Sumatra); P. lamellatus sp.
nov. (Thailand); P. latibasalis sp. nov. (Sri Lanka); P. laticuspis
sp. nov. (Indonesia: Western New Guinea); P. latior sp. nov. (Malaysia;
Brunei); P. limans sp. nov. (Indonesia: Western New Guinea); P.
linguatus sp. nov. (Brunei); P. linguiformis sp. nov. (Laos); P.
loebli sp. nov. (Southwest India); P. longalatus sp. nov. (Indonesia:
Sumatra); P. longastatus sp. nov. (Philippines: Luzon); P.
longilobatus sp. nov. (Philippines: Leyte; Indonesia: Sulawesi); P.
longipalpis sp. nov. (China: Yunnan); P. lunatus sp. nov. (Sri
Lanka); P. monstrans sp. nov. (Nepal; Northeast India); P. peniculatus
sp. nov. (Malaysia;
Indonesia); P. perfodens sp. nov. (Thailand;
Malaysia); P. petiolatus sp. nov. (Myanmar); P. reflexus sp. nov.
(India; Myanmar; China; Thailand; Laos; Vietnam); P. robusticornis sp.
nov. (Papua New Guinea); P. rossii sp. nov. (Thailand; Laos; Cambodia;
Vietnam); P. rostratus sp. nov. (Southwest India); P. rotundatus
sp. nov. (Indonesia: Western New Guinea); P. rubidipennis sp. nov.
(Northwest India); P. schawalleri sp. nov. (Thailand; Laos); P.
schillhammeri sp. nov. (Indonesia:
Sumatra); P. shavrini sp. nov. (Philippines); P. solomonicus sp.
nov. (Solomon Islands); P. spectans sp. nov. (Brunei); P. spiralis
sp. nov. (Sri Lanka); P. substrictus sp. nov. (Solomon
Islands; Papua New Guinea); P. subvexus sp. nov. (Laos); P.
sulawesicus sp. nov. (Indonesia:
Sulawesi); P. tamilus sp. nov. (South
India); P. taurus sp. nov. (Malaysia:
Sabah, Sarawak; Brunei); P. tensissimus sp. nov. (Nepal; India); P.
torifer sp. nov. (Brunei); P. triacer sp. nov. (Papua
New Guinea); P. tricuspis sp. nov. (Indonesia: Western New Guinea; Papua
New Guinea); P. trifidus sp. nov. (Indonesia: Western New Guinea); P.
trigonalis sp. nov. (India; Myanmar; Laos); P. trispinosus sp. nov.
(Brunei); P. unguicus sp. nov. (Philippines: Palawan); P. unguifer
sp. nov. (Indonesia: Sulawesi); P. unicuspis sp. nov. (New Guinea); P.
validicornis sp. nov. (Malaysia: Sarawak), P. vates sp. nov.
(Philippines: Leyte); P. vellicans sp. nov. (South India); P.
vespertilio sp. nov. (Vietnam); P. vesperugo sp. nov. (Thailand;
Cambodia); P. yunnanicus sp. nov. (China:
Yunnan); Pinachenium gen. nov. (type species: Pinophilus longicornis Baudi
di Selve, 1848); Pinobelus gen. nov. (type species: Pinophilus
punctatissimus Sharp, 1889); Pinocavus gen. nov. (type species Pinocavus
excisus sp. nov.; Saudi Arabia); P. volans sp. nov. (South
India); Pinocharis acifera sp. nov. (Laos); P. biacuta sp. nov.
(Myanmar); P. bifida sp. nov. (Thailand; Cambodia); P. burmana sp.
nov. (Myanmar); P. clivis sp. nov. (Thailand; Laos; Cambodia; Vietnam); P.
flagrans sp. nov. (South India; Sri Lanka); P. forans sp. nov. (India;
Sri Lanka); P. furcillata sp. nov. (Laos);
P. migra sp. nov. (Cambodia; Vietnam); P. proiecta sp. nov.
(Thailand; Cambodia); P. unguica sp. nov. (Myanmar); P. velifera
sp. nov. (Thailand; Laos; Cambodia; Vietnam); Pinocolis gen. nov. (type
species: Pinophilus leucopus Kraatz, 1859); P. appendiculatus sp.
nov. (Southwest India); P. asperifrons sp. nov. (Nepal; India: Assam);
P. atrifemoralis sp. nov. (Philippines); P. bifilifer sp. nov. (South
India); P. bisinuatus sp. nov. (China; Thailand; Laos; Vietam); P.
commixtus sp. nov. (Myanmar);
P.
fuscitibialis
sp. nov. (Hong Kong; China); P.
gemellus
sp. nov. (Myanmar);
P. insectus sp. nov. (Indonesia: Kalimantan Selatan); P. mastigans
sp. nov. (Thailand; Malaysia); P. piscator sp. nov. (South India); Pinocrenum
gen. nov. (type species: Pinocrenum dentatum sp. nov.; Nepal); Pinonepalus
aduncus sp. nov. (Malaysia: Perak); P. affimbriatus sp. nov.
(Malaysia: Sabah); P. aquilus sp. nov. (Indonesia:
Kalimantan Tengah); P. atricollis sp. nov. (Philippines: Luzon, Leyte); P.
atricrus sp. nov. (Vietnam; Indonesia: Sumbawa, Lombok); P. auctus
sp. nov. (Indonesia: Sulawesi); P. basoricus sp. nov. (Malaysia:
Kelantan); P. bellipes sp. nov. (Malaysia: Sarawak; Indonesia: Kalimantan
Tengah); P. bicarinatus sp. nov. (Malaysia: Sarawak); P. bicoloripes
sp. nov. (Philippines:
Mindanao); P. biconstrictus sp. nov. (Indonesia: Sumatra); P.
biconvexus sp. nov. (Indonesia:
Sumatra); P. brevilobatus sp. nov. (Indonesia: Sulawesi); P. claripes
sp. nov. (Indonesia: Sumatra); P. coccinus sp. nov. (Indonesia:
Sulawesi); P. contegens sp. nov. (Thailand);
P. contractus sp. nov. (Malaysia); P. corculatus sp. nov. (Indonesia:
Sulawesi); P. cordatus sp. nov. (Indonesia: Sulawesi); P. cybaeus
sp. nov. (Philippines); P. deambulans sp. nov. (Philippines: Mindanao); P.
demorsus sp. nov. (Malaysia: Sabah); P. derectus sp. nov. (Malaysia:
Sarawak); P. disparilis sp. nov. (Malaysia);
P. distortus sp. nov. (Indonesia:
Sumatra); P. districtus sp. nov. (Malaysia: Sabah); P. divexatus
sp. nov. (Indonesia: Sumatra); P. elevatus sp. nov. (Sri Lanka); P.
extensus sp. nov. (Indonesia: Sumatra); P. fissus sp. nov. (India:
Meghalaya); P. fusus sp. nov. (China: Guangxi); P. gilvipes sp.
nov. (Malaysia:
Sabah); P. goanus sp. nov. (South India); P. hamulatus sp. nov. (Indonesia:
Sumatra); P. hebetatus sp. nov. (Indonesia: Sulawesi); P. ingens
sp. nov. (Indonesia: Sulawesi); P. iniquus sp. nov. (Thailand);
P. latilobatus sp. nov. (Thailand); P. ligulatus sp. nov.
(Philippines); P. limus sp. nov. (Malaysia); P. linguifer sp. nov.
(Malaysia: Sarawak); P. luteipennis sp. nov. (Indonesia:
Sumatra); P. luteipes sp. nov. (Philippines: Leyte); P. luzonicus
sp. nov. (Philippines: Luzon); P. mancus sp. nov. (Malaysia); P.
manasicus sp. nov. (India: Assam); P. migrus sp. nov. (Malaysia:
Sabah); P. muluicus sp. nov. (Malaysia: Sarawak); P. muscicola sp.
nov. (Malaysia:
Sarawak); P. nigricrus sp. nov. (Thailand); P. ostentans sp. nov.
(Philippines: Mindanao); P. palatus sp. nov. (Myanmar);
P. parilis sp. nov. (Indonesia: Sumatra); P. pauli sp. nov.
(Indonesia: Bali); P. pellucidus sp. nov. (Malaysia: Sabah, Sarawak); P.
perlucens sp. nov. (Malaysia: Sarawak); P. polisicus sp. nov.
(Philippines: Luzon); P. pravus sp. nov. (Indonesia: Sumatra); P.
procerus sp. nov. (Philippines: Mindanao); P. pungens sp. nov.
(Malaysia); P. retunsus sp. nov. (Indonesia: Sulawesi); P. sabahicus
sp. nov. (Malaysia: Sabah); P. sagittans sp. nov. (Indonesia: Sumatra); P.
sagittarius sp. nov. (Indonesia: Sumatra); P. siporanus sp. nov. (Indonesia:
Metawei Islands); P. subtilior sp. nov. (Indonesia: Sulawesi); P.
taleatus sp. nov. (Thailand); P. tortus sp. nov. (Malaysia:
Selangor); P. translucidus sp. nov. (Malaysia:
Sarawak); P. triquetrus sp. nov. (Laos; Vietnam); P. venetus sp.
nov. (Malaysia:
Sabah); Pinopellis gen. nov. (type species: Pinophilus borneensis
Fauvel, 1895); Pinopellis apona sp. nov. (Philippines: Mindanao; P.
aspera sp. nov. (Borneo: Brunei); P. biexcisa sp. nov. (Myanmar); P.
breviata sp. nov. (Philippines:
Mindanao, Leyte); P. burmana sp. nov. (Myanmar);
P. clavulata sp. nov. (China:
Jiangxi, ?Guizhou); P. laotica sp. nov. (Laos); P. longior sp.
nov. (Malaysia:
Pahang); P. mutabilis sp. nov. (Indonesia:
Sulawesi); P. nepalensis sp. nov. (Nepal);
P. russipes sp. nov. (Philippines: Luzon); P. spiculata sp. nov.
(Philippines: Mindanao); P. tecta sp. nov. (Indonesia: Sulawesi); P.
tensa sp. nov. (Indonesia:
Sulawesi); P. tricarinata sp. nov. (Laos);
P. truncata sp. nov. (Laos); P. turana sp. nov.(India: Meghalaya);
P. volitans sp. nov. (Philippines: Mindanao); P. yunnanica sp.
nov. (China: Yunnan); Pinoplanus gen. nov. (type species: Pinophilus
aegyptius Erichson, 1840); Pinopressus gen. nov. (type species: Pinophilus
complanatus Erichson, 1840); Pinoremus gen. nov. (type species:
Pinoremus cuneatus sp. nov.; Papua New Guinea); Pinorima gen. nov.
(type species: Pinorima spinosa sp. nov.; Thailand; China; Laos;
Cambodia; Vietnam); Pinosenex gen. nov. (type species: Pinosenex
falcatus sp. nov.; South India).
In addition, one genus and four species from other zoogeographic regions are
described: Pinoplanus spathulatus sp. nov. (Kenya; Namibia; Ethiopia); P.
auctior sp. nov. (Tanzania); P. constrictus sp. nov. (South Africa;
Zimbabwe); Novipinophilus gen. nov. (type species: Novipinophilus
caledonicus sp. nov.; New Caledonia).
Five names are revalidated: Pinocolis curtipennis (Bernhauer, 1935)
(previously a synonym of Pinophilus femoratus Schubert, 1911); Pinocolis
insignis (Sharp, 1874) (previously a synonym of Pinophilus javanus
Erichson, 1840); Pinocolis pallipes (Kraatz, 1859) (previously a synonym
of Pinophilus javanus); Pinoplanus densior (Bernhauer, 1935); Pinoplanus
punctulatus (Fauvel, 1905).
One genus-group and 17 species-group synonymies are proposed: Pinophilinus
Eichelbaum, 1908 = Pinogalus Fagel, 1963, syn. nov.; Mimopinophilus
formosae (Bernhauer, 1935) = M. coomani (Bernhauer, 1935), syn. nov.;
Mimopinophilus picticornis (Kraatz, 1859) = M. laocoon (Fernando,
1959), syn. nov.; Mimopinophilus rufipennis (Sharp, 1874) = M.
chinensis (Bernhauer, 1938), syn. nov.; Phinopilus sondaicus
(Bernhauer, 1915) = P. proximatus (Cameron, 1936), syn. nov.; Pinachenium
longicorne (Baudi di Selve, 1848) = P. depressum (Kraatz, 1859), syn.
nov., = P. planum (Kraatz, 1859), syn. nov., = P. verticale
(Fauvel, 1895), syn. nov., = P. wittmeri (Koch, 1934), syn. nov.; Pinobelus
punctatissimus (Sharp, 1889) = P. parapunctatissimus (Li & Wang,
1993), syn. nov., = P. lini (Rougemont, 2017), syn. nov.; Pinocolis
curtipennis (Bernhauer, 1935) = P. assamensis (Coiffait, 1982), syn.
nov.; Pinocolis pallipes (Kraatz, 1859) = P. ferrugineus
(Bernhauer, 1902), syn. nov.; Pinonepalus cassagnaui Coiffait, 1982 =
P. nepalicus (Coiffait, 1982), syn. nov.; Pinonepalus rotundicollis
(Kraatz, 1859) = P. orientalis (Cameron, 1920), syn. nov., = P.
jacobsoni (Cameron, 1930), syn. nov., = P. perakanus (Cameron, 1950),
syn. nov.; Pinopressus complanatus (Erichson, 1840) = P. ruficeps
(Kraatz, 1859), syn. nov. In all, 104 new combinations are proposed, including
42 species distributed in the Afrotropical region: Mimopinophilus formosae
(Bernhauer, 1935), comb. nov. (ex Pinophilus); Mimopinophilus
rufipennis (Sharp, 1874), comb. nov. (ex Pinophilus); Mimopinophilus
sanguinipennis (Bernhauer, 1935), comb. nov. (ex Pinophilus); Neopinophilus
brachypterus (Kraatz, 1859), comb. nov. (ex
Pinophilus); Neopinophilus funestus (Bernhauer, 1902), comb. nov.
(ex Pinophilus); Phinopilus bomfordi (Eppelsheim, 1890), comb.
nov. (ex Pinophilus); Phinopilus brevicornis (Cameron, 1952),
comb. nov. (ex Pinophilus); Phinopilus brevis (Kraatz, 1859),
comb. nov. (ex Pinophilus); Phinopilus ceylonicus (Cameron, 1931),
comb. nov. (ex Pinophilus); Phinopilus drescheri (Cameron, 1936),
comb. nov. (ex Pinophilus); Phinopilus falsus (Cameron, 1940),
comb. nov. (ex Pinophilus); Phinopilus hindostanus (Cameron,
1945), comb. nov. (ex Pinophilus); Phinopilus lewisius (Sharp,
1874), comb. nov. (ex Pinophilus); Phinopilus papuanus (Cameron,
1937), comb. nov. (ex Pinophilus); Phinopilus peninsularis
(Cameron, 1932), comb. nov. (ex Pinophilus); Phinopilus piceus
(Fauvel, 1904), comb. nov. (ex Pinophilus); Phinopilus pilicollis
(Motschulsky, 1858), comb. nov. (ex Pinophilus); Phinopilus
preangeranus (Cameron, 1936), comb. nov. (ex Pinophilus);
Phinopilus sachtlebeni (Bernhauer, 1935), comb. nov. (ex Pinophilus);
Phinopilus sautteri (Bernhauer, 1935), comb. nov. (ex Pinophilus);
Phinopilus senescens (Cameron, 1925), comb. nov. (ex Pinophilus);
Phinopilus sondaicus (Bernhauer, 1915), comb. nov. (ex Pinophilus);
Phinopilus uniformis (Cameron, 1928), comb. nov. (ex
Pinophilus); Pinachenium longicorne (Baudi di Selve, 1949), comb.
nov. (ex Pinophilus); Pinobelus punctatissimus (Sharp, 1889),
comb. nov. (ex Pinophilus); Pinocavus abnormalis (Bernhauer,
1915), comb. nov. (ex Pinophilinus); Pinocavus africanus (Gestro,
1873), comb. nov. (ex Pinophilinus); Pinocavus auberti (Fagel,
1963), comb. nov. (ex Pinophilinus); Pinocavus australis
(Gemminger & Harold, 1868), comb. nov. (ex Pinophilinus); Pinocavus
gestroi (Gridelli, 1928), comb. nov. (ex
Pinophilinus); Pinocavus gibbifrons (Fagel, 1963), comb. nov. (ex Pinophilinus);
Pinocavus grandicollis (Fagel, 1963), comb. nov. (ex Pinophilinus);
Pinocavus itombwensis (Fagel, 1963), comb. nov. (ex Pinophilinus);
Pinocavus kaboboensis (Fagel, 1963), comb. nov. (ex Pinophilinus);
Pinocavus kahuziensis (Fagel, 1963), comb. nov. (ex Pinophilinus);
Pinocavus lamottei (Fagel, 1963), comb. nov. (ex Pinophilinus); Pinocavus
leleupi (Fagel, 1963), comb. nov. (ex Pinophilinus); Pinocavus
luberoensis luberoensis (Fagel, 1963), comb. nov. (ex Pinophilinus); Pinocavus
luberoensis dubius (Fagel, 1963), comb. nov. (ex Pinophilinus); Pinocavus
minutus (Gridelli, 1928), comb. nov. (ex Pinophilinus); Pinocavus
mwengensis (Fagel, 1963), comb. nov. (ex Pinophilinus); Pinocavus
raffrayi (Fagel, 1963), comb. nov. (ex Pinophilinus); Pinocavus
rugegensis (Fagel, 1963), comb. nov. (ex Pinophilinus); Pinocavus
rugosus (Gridelli, 1928), comb. nov. (ex Pinophilinus); Pinocavus
schatzmayri (Koch, 1934), comb. nov. (ex Pinophilinus); Pinocavus
sjostedti (Eichelbaum, 1908), comb. nov. (ex Pinophilinus); Pinocavus
socotranus (Assing, 2012), comb. nov. (ex Pinophilinus); Pinocavus
somalicus (Fagel, 1963), comb. nov. (ex Pinophilinus); Pinocavus
spinosus (Assing, 2013), comb. nov. (ex Pinophilinus); Pinocavus
strictipennis (Fagel, 1963), comb. nov. (ex Pinophilinus); Pinocavus
tshuapaensis (Fagel, 1963), comb. nov. (ex Pinophilinus); Pinocharis
genalis (Fauvel, 1895), comb. nov. (ex Pinophilus); Pinocharis
rufoferruginea (Cameron, 1936), comb. nov. (ex Pinophilus); Pinocolis
chapmani (Bernhauer, 1935), comb. nov. (ex Pinophilus); Pinocolis
curtipennis (Bernhauer, 1935), comb. nov. (ex Pinophilus);
Pinocolis femoratus (Schubert, 1911), comb. nov. (ex Pinophilus); Pinocolis
insignis (Sharp, 1874), comb. nov. (ex Pinophilus); Pinocolis
javanus (Erichson, 1840), comb. nov. (ex Pinophilus); Pinocolis
leucopus (Kraatz, 1859), comb. nov. (ex Pinophilus); Pinocolis
malabarensis (Cameron, 1931), comb. nov. (ex Pinophilus);
Pinocolis mixtus (Cameron, 1914), comb. nov. (ex Pinophilus);
Pinocolis pallipes (Kraatz, 1859), comb. nov. (ex Pinophilus);
Pinocolis variipes (Fauvel, 1895), comb. nov. (ex Pinophilus);
Pinocolis vicinus (Cameron, 1931), comb. nov. (ex Pinophilus);
Pinonepalus amicus (Bernhauer, 1915), comb. nov. (ex Pinophilus);
Pinonepalus beccarii (Fauvel, 1895), comb. nov. (ex Pinophilus);
Pinonepalus bettotanus (Cameron, 1930), comb. nov. (ex Pinophilus);
Pinonepalus brunneipennis (Cameron, 1932), comb. nov. (ex Pinophilus);
Pinonepalus cameroni (Scheerpeltz, 1933), comb. nov. (ex Pinophilus);
Pinonepalus eppelsheimi (Bernhauer, 1904), comb. nov. (ex Pinophilus);
Pinonepalus melanocephalus (Motschulsky, 1858), comb. nov. (ex Pinophilus);
Pinonepalus melanomerus (Kraatz, 1859), comb. nov. (ex Pinophilus); Pinonepalus
parvus (Bernhauer, 1914), comb. nov. (ex Pinophilus); Pinonepalus
philippinus (Bernhauer, 1915), comb. nov. (ex Pinophilus);
Pinonepalus rotundicollis (Kraatz, 1859), comb. nov. (ex Pinophilus);
Pinonepalus selangorensis (Cameron, 1950), comb. nov. (ex Pinophilus);
Pinonepalus separandus (Cameron, 1928), comb. nov. (ex Pinophilus);
Pinonepalus thoracicus (Fauvel, 1895), comb. nov. (ex Pinophilus);
Pinopellis borneensis (Fauvel, 1895), comb. nov. (ex Pinophilus);
Pinopellis coarcticollis (Cameron, 1941), comb. nov. (ex Pinophilus);
Pinopellis nigripes (Cameron, 1914), comb. nov. (ex Pinophilus); Pinophilinus
anosyanus (Lecoq, 1986), comb. nov. (ex Pinogalus); Pinophilinus
betschi (Jarrige, 1978), comb. nov. (ex Pinogalus); Pinophilinus
brunneorufus (Bernhauer, 1937), comb. nov. (ex Pinogalus); Pinophilinus
cameroni (Fagel, 1963), comb. nov. (ex Pinogalus); Pinophilinus
clementi (Lecoq, 1986), comb. nov. (ex Pinogalus); Pinophilinus
cordicollis (Jarrige, 1978), comb. nov. (ex Pinogalus); Pinophilinus
daressalamensis (Bernhauer, 1937), comb. nov. (ex Pinogalus); Pinophilinus
delkeskampi (Fagel, 1963), comb. nov. (ex Pinogalus); Pinophilinus
fauveli (Schubert, 1902), comb. nov. (ex Pinogalus); Pinophilinus
levasseuri (Lecoq, 1986), comb. nov. (ex Pinogalus); Pinophilinus
mantasoae (Lecoq, 1986), comb. nov. (ex Pinogalus); Pinophilinus
matumbianus (Bernhauer, 1937), comb. nov. (ex Pinogalus); Pinophilinus
micropterus (Bernhauer, 1937), comb. nov. (ex Pinogalus); Pinophilinus
peyrierasi (Lecoq, 1986), comb. nov. (ex Pinogalus); Pinophilinus
rufus (Lecoq, 1986), comb. nov. (ex Pinogalus); Pinophilinus
schuberti (Fauvel, 1905), comb. nov. (ex Pinogalus); Pinophilinus
zambezianus (Fagel, 1963), comb. nov. (ex Pinogalus); Pinoplanus
aegyptius (Erichson, 1840), comb. nov. (ex Pinophilus); Pinoplanus
grandis (Reitter, 1906), comb. nov. (ex Pinophilus); Pinoplanus
densior (Bernhauer, 1935), comb. nov.; Pinoplanus punctulatus
(Fauvel, 1905), comb. nov.; Pinopressus complanatus (Erichson, 1840),
comb. nov. (ex Pinophilus); Pinopressus indicus (Cameron, 1931),
comb. nov. (ex
Pinophilus).
Mimopinophilus sanguinipennis
(Bernhauer, 1935), previously a subspecies of M. formosae (Bernhauer,
1935) is regarded as a distinct species.
A neotype is designated for Pinophilus siculus Kraatz, 1857. Lectotypes
are designated for Pinophilus fossor Wollaston, 1868, P. zarudnyi
Semenov, 1909, P. formosae Bernhauer, 1935, P. coomani Bernhauer,
1935, P. picticornis Kraatz, 1859, P. rufipennis Sharp, 1874, P.
chinensis Bernhauer, 1938, P. sanguinipennis Bernhauer, 1935, P.
punctatissimus Sharp, 1889, P. pallipes Kraatz, 1859, P. chapmani
Bernhauer, 1935, P. variipes Fauvel, 1895, P. vicinus Cameron,
1931, P. malabarensis Cameron, 1931, P. curtipennis Bernhauer,
1935, P. mixtus Cameron, 1914, Pinophilus ruficeps Kraatz, 1859, P.
indicus Cameron, 1931, P. depressus Kraatz, 1859, P. wittmeri
Koch, 1934, P. brevicollis Erichson, 1840, P. bomfordi Eppelsheim,
1890, P. sachtlebeni Bernhauer, 1935, P. falsus Cameron, 1940, P.
sautteri Bernhauer, 1935, P. ceylonicus Cameron, 1931, P. papuanus
Cameron, 1937, P. uniformis Cameron, 1928, P. eppelsheimi
Bernhauer, 1904, P. melanomerus Kraatz, 1859, P. rotundicollis
Kraatz, 1859, P. amicus Bernhauer, 1915, P. philippinus Bernhauer,
1915, P. speculifrons Cameron, 1914, P. borneensis Fauvel, 1895, P.
nigripes Cameron, 1914, P. coarcticollis Cameron, 1941, and P.
genalis Fauvel, 1895,
Numerous examined species are represented only by females and consequently not
named. This and the general rarity of records suggest that a substantial number
of species remains to be discovered and described in the future.
A comprehensive catalogue and keys to the genera and species of Pinophilina of
the Palaearctic and Oriental regions are provided.
Aside from a derived aedeagal morphology, the genera represented in the study
region share two evident synapomorphies: fused abdominal tergites IX and X and
strongly dilated, compact protarsomeres IIV (usually broader than combined
length). The sole exception is Pinopressus, a genus characterised by an
aedeagus of primitive condition, unfused tergites IX and X, and moderately
dilated (not compact) protarsomeres IIV. These observations suggest that Pinopressus
forms a basal lineage within Pinophilina. At the generic level, there is
currently no overlap between the fauna of the study region and that of the New
World. On the other hand, many genera distributed in the Palaearctic and
Oriental regions are present also in sub-Saharan Africa and/or Australia.
Species diversity is highest in the warmer climates of the Oriental region,
particularly in continental Southeast Asia and the Sunda Islands. With few
exceptions (records with specified localities unknown), the distributions of all
the species in the study region are mapped.
Rather little is known regarding the natural history of Pinophilina. Most of the
species were found at low to intermediate altitudes, few at high elevations (up
to more than 3000 m). A substantial proportion of the species and specimens
examined was collected primarily or exclusively at light, mostly close to lakes
or rivers and streams. A number of species was exclusively extracted from litter
and soil of forests, partly also from dead wood (bark, logs). The available
ecological data and the general rarity of records suggest that the reproduction
habitats are of a cryptic, partly probably subterranean nature.